Research Article |
Corresponding author: Ryo Ogawa ( ailmasbl0854@yahoo.co.jp ) Academic editor: James Liebherr
© 2016 Ryo Ogawa, Ivan Löbl.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Ogawa R, Löbl I (2016) A review of the genus Xotidium Löbl, 1992 (Coleoptera, Staphylinidae, Scaphidiinae), with descriptions of five new species. Deutsche Entomologische Zeitschrift 63(1): 155-169. https://doi.org/10.3897/dez.63.8386
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In the present study, we have reviewed all species of the genus Xotidium Löbl, 1992, as well as five new species from Borneo, Malaysia, and Sulawesi, Indonesia, namely, Xotidium flagellum sp. n., X. heissi sp. n., X. meridionale sp. n., X. smetanai sp. n., X. tarantulatum sp. n. A key to all Xotidium species is provided. The female genitalic characters of Xotidium are illustrated and described for the first time.
Shining fungus beetle, Baeocera group, female genitalia, Borneo, Sulawesi, key to species
The genus Xotidium Löbl, 1992, currently comprises seven species from the subtropical and tropical regions of Mauritius, Sri Lanka, the Himalayas, the Lesser Sunda Islands, the Philippines, and Eastern Australia. Members of this genus have narrow body with approximate mesocoxae and metacoxae. They appear to be related to the members of Scaphoxium Löbl, 1979 and Toxidium LeConte, 1860, belonging to the Toxidium group because of the shared approximate mesocoxae and metacoxae. However, this genus was included in the Baeocera group by
In the present study, we have reviewed all the members of Xotidium and describe five new species collected from Borneo, Malaysia, and Sulawesi, Indonesia, and present the first reported observations of the female genitalia in Xotidium. Moreover, we provide a key to all Xotidium species, including these new species.
The examined samples were collected by the first author and procured from the collections deposited at the
The abbreviations used are as follows: EL, length of elytra from base of pronotum to apex of elytra; EW, maximum width of elytra; HW, maximum width of head including eye; ID, interocular distance; PL, maximum length of pronotum; PW, maximum width of pronotum;
Xotidium Löbl, 1992: 573; type species: Xotidium uniforme Löbl, 1992.
Body narrow, usually 1.8 times as long as wide. Maxillary palpomere IV aciculate with some macrosetae (Fig.
The shape of the labral apex in this genus is possibly species specific (Fig.
Xotidium bolmarum Löbl, 2015: 119.
Paratypes, 1♂4♀, INDONESIA, LOMBOK IS., SENARO, N slope of Rinjani, 2–5. Feb. 1994, Bolm lgt. 1100m. (
Ovipositor simple: gonostylus with a long apical seta, almost as long as wide: distal gonocoxites elongated (Fig.
(n = 4). Length (PL+EL): 1.40–1.48 mm, PW: 0.77–0.79 mm, EW: 0.83–0.87 mm, HW: 0.31–0.35 mm, ID: 0.10–0.13 mm, PL/PW: 0.71–0.78, EL/EW: 1.00–1.05. Approximate ratio of each antennal segment in length (width) (n = 1): II 1.1 (0.5) : III 1.0 (0.2) : IV 1.1 (0.2) : V 1.3 (0.2) : VI 1.4 (0.2) : VII 1.4 (0.3) : VIII 1.4 (0.2) : IX 1.7 (0.3) : X 1.3 (0.4) : XI 1.7 (0.5).
(see
Toxidium
mauritianum
Vinson, 1943: 201. Transferred:
2♂1♀, Mauritius, Le Pouce, 18. XI. 1943, J. Vinson. (
Ovipositor simple: gonostylus short, longer than wide, with a long apical seta: distal gonocoxites elongated and slightly robust (Fig.
(n = 3). Length (PL+EL): 1.34–1.51 mm, PW: 0.72–0.79 mm, EW: 0.75–0.85 mm, HW: 0.32–0.34 mm, ID: 0.11–0.13 mm, PL/PW: 0.71–0.74, EL/EW: 1.08–1.11. Approximate ratio of each antennal segment in length (width) (n = 1): II 1.5 (0.5) : III 1.0 (0.3) : IV 1.0 (0.3) : V 1.0 (0.3) : VI 1.1 (0.3) : VII 1.5 (0.4) : VIII 1.1 (0.3) : IX 1.5 (0.5) : X 1.5 (0.6) : XI 1.8 (0.7).
(referred to
Toxidium
montanum
Löbl, 1971: 1000. Transferred:
Holotype, 1♂, CEYLAN Central, Nuwera Eliya, 1950m, 29. I. 1970, Mussard Besuchet Löbl,
Paratypes, 4♂, same data as holotype; 2♂1♀, same data as holotype, but 15. II. 1970. (
1♀, SRI LANKA, 29. iv.–1. v., Idalgashinna, 1400m, 25km SW Badulla, Z. Kejval leg. 1994 (
Ovipositor simple: gonostylus with a long apical seta, distinctly longer than wide; distal gonocoxites elongated (Fig.
(n = 6). Length (PL+EL): 1.38–1.48 mm, PW: 0.73–0.83 mm, EW: 0.79–0.87 mm, HW: 0.32–0.36 mm, ID: 0.13–0.14 mm, PL/PW: 0.64–0.75, EL/EW: 1.00–1.10. Approximate ratio of each antennal segment in length (width) (n = 1): II 1.4 (0.5) : III 1.0 (0.3) : IV 1.1 (0.3) : V 1.3 (0.3) : VI 1.3 (0.3) : VII 1.6 (0.5) : VIII 1.3 (0.4) : IX 1.9 (0.6) : X 1.8 (0.6) : XI 1.9 (0.8).
(see
This species is variable in the color of elytra as shown in Fig.
Toxidium
notatum
Löbl, 1977: 65. Transferred:
Holotype, 1♂, Acacia Plat. N. S. W., J. Armstrong, W. Steel coll., B. M. 1969-552 (NHM).
Paratype, 1♂, N. S. Wales, Acacia Plat, J. Armstrong (
3♂, Queensland, Umg, Bisbane, Wachtel. V. 1981; 1♂1♀, Monsildale Via Kilcay, S. E. Qld., 22. IV. 1963, Monteith; 1♀, N. S. W. Sydeny, Oxford falls, litter + bails, Au: 33, 29. X. 1982, Endrödy-Younga (
Ovipositor simple: gonostylus with a long apical seta, almost as long as wide: distal gonocoxites elongated (Fig.
(n = 5). Length (PL+EL): 1.19–1.38 mm, PW: 0.62–0.68 mm, EW: 0.66–0.74 mm, HW: 0.32–0.36 mm, ID: 0.09–0.13 mm, PL/PW: 0.67–0.74, EL/EW: 1.08–1.24. Approximate ratio of each antennal segment in length (width) (n = 1): II 1.2 (0.5) : III 1.0 (0.3) : IV 1.0 (0.3) : V 1.2 (0.3) : VI 1.3 (0.3) : VII 1.5 (0.4) : VIII 1.4 (0.3) : IX 1.6 (0.5) : X 1.5 (0.5) : XI 2.0 (0.6).
(see
The male genitalia have a long sclerite (Fig.
Toxidium
pygmaeum
Löbl, 1971: 998. Transferred:
Holotype, 1♂, CEYLAN Central, Kandy, 700 m, 16. II. 1970, Mussard Besuchet Löbl,
Paratypes, 9♂2♀, same data as holotype, but 14. II. 1970 (
Ovipositor simple: gonostylus with a long apical seta, distinctly longer than wide; distal gonocoxites elongated (Fig.
(n = 5). Length (PL+EL): 1.11–1.19 mm, PW: 0.66–0.68 mm, EW: 0.68–0.74 mm, HW: 0.26–0.30 mm, ID: 0.09–0.11 mm, PL/PW: 0.66–0.69, EL/EW: 0.97–1.03. Approximate ratio of each antennal segment in length (width) (n = 1): II 1.3 (0.7) : III 1.0 (0.4) : IV 1.1 (0.3) : V 1.2 (0.4) : VI 1.3 (0.4) : VII 1.5 (0.5) : VIII 1.3 (0.4) : IX 1.7 (0.6) : X 1.3 (0.6) : XI 1.7 (0.7).
(see
Xotidium tubuliferum Löbl, 2011: 720.
1♂, Tarragona, Leyte I. Philippine islds VIII: II: 1945 / Col. & pres. by C. L. Remington /in rotting burned log. [pronotum and head missing] (
(referred to
Xotidium uniforme Löbl, 1992: 575.
Holotype, 1♂, E. NEPAL: KOSI, Val. Induwa Kola, 2100 m, 17. IV. 1984, Löbl – Smetana (
Paratypes, 1♀, INDE Kumaon (UP), Rangarh nr. 6b, ewu. 2000m, I. Löbl, 9. X. 1979; 2♂, INDE: HTM. PM. PR., 10km, NW Sarahan, 1700m, 7. X. 1988, S. Vit; 3♂1♀, 414, Sankhua Sabha Distr., Arun Valley, Chichila, 1900-2000m, Quercus forest, bushes near village, 18-20. VI. 1988, J. Martens & W. Schawaller (
Ovipositor simple: gonostylus with a very long apical seta, slightly longer than wide: distal gonocoxites elongated (Fig.
(n = 5). Length (PL+EL): 1.58–1.63 mm, PW: 0.85–0.88mm, EW: 0.92–0.96 mm, HW: 0.36–0.38 mm, ID: 0.10–0.14 mm, PL/PW: 0.71–0.75, EL/EW: 1.02–1.08. Approximate ratio of each antennal segment in length (width) (n = 1): II 1.4 (0.6) : III 1.0 (0.2) : IV 1.2 (0.3) : V 1.3 (0.2) : VI 1.3 (0.3) : VII 1.6 (0.4) : VIII 1.4 (0.3) : IX 1.8 (0.4) : X 1.5 (0.5) : XI 1.6 (0.5).
(see
Holotype, 1♂, Borneo Sabah, Mt. Kinabalu N. P. summit tr. Pondok Lowii, 2300–2400 m, 28. IV. 1987, A. Smetana,
The species epithet refers to the presence of a flagellum in the internal sac.
(male). Dorsal and ventral surface almost reddish-brown (Fig.
Head with eye width almost the same as interocular distance. Punctuation sparse and fine.
Pronotum wider than long. Punctuation sparse and fine, as on head. Scutellum with exposed apex.
Elytra as long as wide, widest at basal fourth, lateral margins sharply narrowed apically, minutely serrate at inner part of posterior margin. Punctuation fine and sparse as on pronotum. Sutural striae extending outwards along basal margin to form basal striae, reaching humeral area and joined with lateral striae.
Hypomeron and lateral portion of mesoventrite smooth. Lateral portion of metaventrite finely and sparsely punctate. Metanepisternum about four times as long as wide, without longitudinal line. Mesepimeron almost as long as wide. Metacoxa about six times as wide as metacoxal process. Ventrite I sparsely and finely punctate.
Protarsomere V about three times as long as each I–IV. Mesotarsomere I about three times as long as each II–IV; V about 2.5 times as long as each II–IV. Metatarsomere I about twice as long as each II and III; each II and III about 1.5 times as long as IV; V about twice as long as IV.
Protarsomeres I–III possibly with tenent setae, but invisible at ×80 magnification, not enlarged. Aedeagus about 0.52 mm long; parameres symmetrical, with moderately widened apical section much longer than third of total parameral length; internal sac with a sinuate and curved flagellum, widened basally and apically (Fig.
Female. Unknown.
(n = 1). Length (PL+EL): 1.23 mm, PW: 0.65 mm, EW: 0.69 mm, HW: 0.30 mm, ID: 0.13 mm, PL/PW: 0.62, EL/EW: 1.19. Approximate ratio of each antennal segment in length (width) (n = 1): II 1.4 (0.7) : III 1.0 (0.3) : IV 1.1 (0.2) : V 1.3 (0.3) : VI 1.5 (0.3) : VII 1.7 (0.4) : VIII 1.6 (0.3) : IX 1.9 (0.5) : X 1.7 (0.5) : XI 1.9 (0.6).
East Malaysia: Sabah.
This species is very similar to X. pygmaeum (Löbl) in the smaller body, but may be easily distinguished by the very long flagellar sclerite of male genitalia.
Holotype, 1♂, Brunei/Temburong, Kuala Belalong FSC, 60–300 m, 16–20. IV. 1993, mixed Dipterc. for., E. Heiss,
Paratype, 1♂, SABAH: Porig Hot, Springs, 500 m, 6. V. 1987, Burckhardt – Löbl,
This species epithet is named in honour of the collector of the holotype, E. Heiss from Innsbruck, Austria.
(male). Dorsal and ventral surface almost reddish-brown (Fig.
Head with eye width almost the same as interocular distance. Punctuation sparse and fine.
Pronotum wider than long. Punctuation sparse and fine, as on head. Scutellum almost as long as wide, with exposed apex.
Elytra almost as long as wide, widest at basal fourth, lateral margins sharply narrowed apically, minutely serrate at inner part of posterior margin. Punctuation fine and sparse as on pronotum. Sutural striae extending outwards along basal margin to form basal striae, reaching humeral area and joined with lateral striae.
Hypomeron and lateral portion of mesoventrite smooth. Lateral portion of metaventrite finely and sparsely punctate. Metanepisternum about four times as long as wide, without longitudinal line. Metepimeron almost as long as wide. Metacoxa about six times as wide as metacoxal process. Ventrite I sparsely and finely punctate.
Protarsomere V about 2.5 times as long as each I–IV. Mesotarsomere I about 2.5 times as long as each II–IV; V about twice as long as IV. Metatarsomere I about three times as long as each II–IV; V about 1.5 times as long as each II–IV.
Protarsomeres I–III possibly with tenent setae, but invisible at ×80 magnification, not enlarged. Aedeagus 0.54 mm long; parameres symmetrical, with strongly widened apical section longer than third of total parameral length; internal sac with a sinuate flagellar sclerite, basally widened and bent (Fig.
Female. Unknown.
(n = 2). Length (PL+EL): 1.33–1.38 mm, PW: 0.73 mm, EW: 0.75–0.78 mm, HW: 0.33–0.35 mm, ID: 0.13 mm, PL/PW: 0.69–0.72, EL/EW: 1.10. Approximate ratio of each antennal segment in length (width) (n = 1): II 1.2 (0.4) : III 1.0 (0.3) : IV 1.4 (0.3) : V 1.6 (0.3) : VI 1.8 (0.2) : VII 2.0 (0.3) : VIII 1.8 (0.3) : IX 2.1 (0.5) : X 1.9 (0.5) : XI 2.3 (0.5).
Brunei; East Malaysia: Sabah.
This species is very similar to X. bolmarum Löbl in the reddish body and the body size, but may be easily distinguished by the strongly widened apical portion of parameres and the sinuate sclerite of male genitalia.
Holotype, 1♂, Mt. Lompobatang, Malino, S. Sulawesi, alt. ca. 1700m, 5°23’44.20”N, 119°55’22.27”E, 20. I. 2011, R. Ogawa leg. (MZBI).
The species epithet is from the Latin meridionale meaning southern, referring to Lompobatang located at the southernmost Sulawesi.
(male). Dorsal and ventral surface almost dark reddish-brown to brown (Fig.
Head with eye width almost the same as interocular distance. Punctuation sparse and fine.
Pronotum wider than long. Punctuation sparse and fine, as on head. Scutellum with slightly exposed apex.
Elytra longer than wide, widest at basal sixth, lateral margins sharply narrowed apically, minutely serrate at inner part of posterior margin. Punctuation fine and sparse as on pronotum. Sutural striae interrupted at basal third.
Hypomeron and lateral portion of mesoventrite smooth. Lateral portion of metaventrite finely and sparsely punctate. Metanepisternum about four times as long as wide, without longitudinal line. Mesepimeron concealed. Metacoxa about six times as wide as metacoxal process. Ventrite I sparsely and finely punctate.
Protarsomere V about three times as long as each I–IV. Mesotarsomere I about twice as long as each II and III; V about twice as long as IV. Metatarsomere I about 2.5 times as long as each II and III; V about three times as long as IV.
Protarsomeres I–III possibly with tenent setae, but invisible at ×80 magnification, not enlarged. Aedeagus 0.54 mm long; parameres symmetrical, with moderately widened apical section shorter than third of total parameral length; internal sac with a sinuate and very long flagellum (longer than aedeagus), basally widened and bent (Fig.
Female. Unknown.
(n = 1). Length (PL+EL): 1.43 mm, PW: 0.81 mm, EW: 0.87 mm, HW: 0.34 mm, ID: 0.11mm, PL/PW: 0.72, EL/EW: 0.98. Approximate ratio of each antennal segment in length (width) (n = 1): II 1.5 (0.6) : III 1.0 (0.3) : IV 1.2 (0.3) : V 1.3 (0.3) : VI 1.5 (0.3) : VII 1.8 (0.4) : VIII 1.7 (0.3) : IX 2.0 (0.5) : X 1.9 (0.5) : XI 2.1 (0.7).
Indonesia: southern Sulawesi.
This species is similar to X. smetanai sp. n. in lacking sutural striae and X. tubuliferum Löbl in the long sclerite as flagellum of male genitalia, but it is easily distinguished from them by the body color and the widened apical section of parameres.
Holotype, 1♂, Borneo Sabah, Mt. Kinabalu Nat. Pk. HQ Silau-Silau, Tr. 1550 m, 2. IV. 1988, [B171], A. Smetana,
This species epithet is in honour of the collector of its holotype, A. Smetana, Ottawa Canada.
Dorsal and ventral surface almost brown (Fig.
Head with eye width almost the same as interocular distance. Punctuation sparse and fine.
Pronotum wider than long. Punctuation sparse and fine, as on head. Scutellum with slightly exposed apex.
Elytra longer than wide, widest at basal sixth, lateral margins sharply narrowed apically, minutely serrate at inner part of posterior margin. Punctuation fine and sparse as on pronotum. Sutural striae interrupted at basal third.
Hypomeron and lateral portion of mesoventrite smooth. Lateral portion of metaventrite finely and sparsely punctate. Metanepisternum about four times as long as wide, without longitudinal line. Mesepimeron almost as long as wide. Metacoxa about six times as wide as metacoxal process. Ventrite I sparsely and finely punctate.
Protarsomere V 2.5 times as long as each I–IV. Mesotarsomere I about twice as long as II; II 1.2 times as long as each III and IV; V 1.5 times as long as each III and IV. Metatarsomere I about three times as long as each II and III; each II and III 1.1 times as long as IV; V about twice as long as IV.
Protarsomeres I–III possibly with tenent setae, but invisible at ×80 magnification, not enlarged. Aedeagus about 0.5 mm long; parameres symmetrical, with moderately widened apical section slightly longer than third of total parameral length; internal sac with straight, evenly wide flagellar sclerite (Fig.
Female. Unknown.
(n = 1). Length (PL+EL): 1.53 mm, PW: 0.85 mm, EW: 0.88 mm, HW: 0.35 mm, ID: 0.15 mm, PL/PW: 0.62, EL/EW: 1.14. Approximate ratio of each antennal segment in length (width) (n = 1); II 1.5 (0.5) : III 1.0 (0.2) : IV 1.1 (0.3) : V 1.3 (0.2) : VI 1.3 (0.2) : VII 1.7 (0.3) : VIII 1.5 (0.3) : IX 1.8 (0.3): X 1.7 (0.4) [XI deformed].
East Malaysia: Sabah.
This species is similar to X. mauritianum (Vinson) and X. meridionale sp. n. in the reduced sutural striae, but may be easily distinguished from them by the simple sclerite of the male genitalia.
Holotype, 1♂, Mt Tilongkabila (Gunung Tilongkabila), N. Sulawesi, alt. ca. 1300m, 0°35’18.14”N, 123°13’22.71”E, 10. VI. 2012, R. Ogawa leg. (MZBI).
Paratypes, 1♂1♀, same data as holotype (
The species epithet is derived from Mapala Tarantula, the climbing club of Gorontalo State University.
Dorsal and ventral surface almost black to dark reddish-brown (Fig.
Head with eye width almost the same as interocular distance. Punctuation sparse and fine.
Pronotum slightly wider than long, with an anterior bead. Punctuation sparse and fine, as on head. Scutellum almost as long as wide, with exposed apex.
Elytra almost as long as wide, widest at basal sixth, lateral margins sharply narrowed apically, minutely serrate at inner part of posterior margin. Punctuation fine and sparse as on pronotum. Sutural striae extending outwards along basal margin to form basal striae, reaching humeral area and joined with lateral striae.
Hypomeron and lateral portion of mesoventrite smooth. Lateral portion of metaventrite finely and sparsely punctate. Metanepisternum about four times as long as wide, without longitudinal line. Mesepimeron almost as long as wide. Metacoxa about six times as wide as metacoxal process. Ventrite I sparsely and finely punctate, with microsculpture.
Protarsomere I twice as long as each II–IV; V 2.5 times as long as each II–IV. Mesotarsomere I 2.5 times as long as II; II 1.2 times as long as each III and IV; V 2.5 times as long as each III and IV. Metatarsomere I about three times as long as each II and III; each II and III 1.1 times as long as IV; V about twice as long as IV.
Male. Protarsomeres I–III possibly with tenent setae, but invisible at ×80 magnification, not enlarged. Aedeagus about 0.44 mm long; parameres symmetrical, with weakly widened apical section as long as fourth of total parameral length; internal sac with a sclerite as flagellum, straight, evenly wide, sometimes bent at base.
Female. Protarsomeres I–III not enlarged. Ovipositor simple: gonostylus with a long apical seta, distinctly longer than wide; distal gonocoxites apparently elongated (Fig.
(n = 9). Length (PL+EL): 1.32–1.43 mm, PW: 0.75–0.79 mm, EW: 0.79–0.89 mm, HW: 0.32–0.36 mm, ID: 0.09–0.13 mm, PL/PW: 0.69–0.76, EL/EW: 0.89–1.02. Approximate ratio of each antennal segment in length (width) (n = 1): II 1.3 (0.7) : III 1.0 (0.3) : IV 1.1 (0.2) : V 1.4 (0.3) : VI 1.6 (0.3) : VII 1.7 (0.4) : VIII 1.6 (0.2) : IX 1.9 (0.4) : X 1.7 (0.4) : XI 2.2 (0.5).
Indonesia: northern Sulawesi.
This species is similar to X. bolmarum Löbl from Lombok in its color and body size, but may be easily distinguished by the straight and more elongate sclerite in the internal sac.
1 | Elytra without basal striae, sutural striae shortened | 2 |
– | Elytra with basal striae, joined to sutural striae | 4 |
2 | Body blackish. Scutellum concealed. Antennomere VIII almost as long as III. Parameres evenly widened. Internal sac without sclerites, slightly sclerotized | X. mauritianum (Vinson) |
– | Body brownish. Scutellum slightly exposed. Antennomere VIII more than 1.5 times as long as III. Parameres notably widened at apical portion. Internal sac with a sclerite | 3 |
3 | Widened apical section of parameres much shorter than a third of total parameral length. Internal sac of aedeagus with a very long flagellum | X. meridionale sp. n. |
– | Widened apical section of parameres much longer than a third of total parameral length. Internal sac of aedeagus with a straight sclerite | X. smetanai sp. n. |
4 | Elytra with distinctive bicolour pattern | 5 |
– | Elytra uniformly reddish-brown to black | 6 |
5 | Elytra dark, each with light transverse fascia situated in basal half of elytron and light apical part. Pronotum entirely dark | X. montanum (Löbl) |
– | Elytra light, each darkened along basal and apical margins, usually also darkened along sutural margin. Pronotum light, usually with dark transverse fascia | X. notatum (Löbl) |
6 | Body length (PL+EL) 1.10–1.25 mm | 7 |
– | Body length (PL+EL) 1.35–1.65 mm | 8 |
7 | Antennomere X distinctly more than 1.5 times as long as III. Mesotarsomeres II and III each about three times as long as I. Metatarsomeres II and III each about twice as long as I. Internal sac of aedeagus with evenly thick and sinuate sclerite | X. pygmaeum (Löbl) |
– | Antennomere X about 1.5 times as long as III. Mesotarsomeres II and III each about twice as long as I. Metatarsomeres II and III each three times as long as I. Internal sac of aedeagus with flagellum curved, sinuate, basally and apically thickened | X. flagellum sp. n. |
8 | Widened apical section of parameres about as long as half of total parameral length. Internal sac of aedeagus with very long, weekly sclerotized and simple tube | X. tubuliferum Löbl |
– | Widened apical section of parameres shorter than half of total parameral length | 9 |
9 | Widened apical section of parameres much longer than third of total parameral length | 10 |
– | Widened apical section of parameres much shorter than third of total parameral length | 11 |
10 | Body length (PL+EL) 1.58–1.63 mm. Antennomere XI less than twice as long as III. Internal sac of aedeagus with an U-shaped flagellum, basally thickened | X. uniforme Löbl |
– | Body length (PL+EL) 1.33–1.38 mm. Antennomere XI more than twice as long as III. Internal sac of aedeagus with a sinuate flagellar sclerite, basally thickened and bent | X. heissi sp. n. |
11 | Antennomere XI more than twice as long as III. Internal sac of aedeagus with straight, evenly thickened flagellum | X. tarantulatum sp. n. |
– | Antennomere XI less than twice as long as III. Internal sac of aedeagus with short basal sclerites followed by long membranous tube | X. bolmarum Löbl |
We wish to express our sincere gratitude to Roger Booth (NHM, London) for the loan of specimens collected by the Project Wallace Expedition and to Giulio Cuccodoro (