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<article xmlns:tp="http://www.plazi.org/taxpub" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:mml="http://www.w3.org/1998/Math/MathML" article-type="review-article" dtd-version="3.0" xml:lang="en">
  <front>
    <journal-meta>
      <journal-id journal-id-type="publisher-id">14</journal-id>
      <journal-id journal-id-type="index">urn:lsid:arphahub.com:pub:69C4EDC7-0CE2-5AB0-81F2-EA3B3CC9D365</journal-id>
      <journal-id journal-id-type="aggregator">urn:lsid:zoobank.org:pub:777322AB-37CC-4A41-B3C2-83A76AAD4F84</journal-id>
      <journal-title-group>
        <journal-title xml:lang="en">Deutsche Entomologische Zeitschrift</journal-title>
        <abbrev-journal-title xml:lang="en">DEZ</abbrev-journal-title>
      </journal-title-group>
      <issn pub-type="ppub">1435-1951</issn>
      <issn pub-type="epub">1860-1324</issn>
      <publisher>
        <publisher-name>Pensoft Publishers</publisher-name>
      </publisher>
    </journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.3897/dez.61.8793</article-id>
      <article-id pub-id-type="publisher-id">4293</article-id>
      <article-categories>
        <subj-group subj-group-type="heading">
          <subject>Review Article</subject>
        </subj-group>
        <subj-group subj-group-type="scientific_subject">
          <subject>Taxonomy</subject>
        </subj-group>
        <subj-group subj-group-type="geographical_area">
          <subject>Asia</subject>
          <subject>Far East</subject>
        </subj-group>
      </article-categories>
      <title-group>
        <article-title>Species of the pleasing lacewing genus <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> Rambur (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order">Neuroptera</tp:taxon-name-part></tp:taxon-name>, <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="family">Dilaridae</tp:taxon-name-part></tp:taxon-name>) from islands of East Asia</article-title>
      </title-group>
      <contrib-group content-type="authors">
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Zhang</surname>
            <given-names>Wei</given-names>
          </name>
          <xref ref-type="aff" rid="A1">1</xref>
        </contrib>
        <contrib contrib-type="author" corresp="yes">
          <name name-style="western">
            <surname>Liu</surname>
            <given-names>Xingyue</given-names>
          </name>
          <email xlink:type="simple">xingyue_liu@yahoo.com</email>
          <uri content-type="orcid">https://orcid.org/0000-0002-9168-0659</uri>
          <xref ref-type="aff" rid="A1">1</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Aspöck</surname>
            <given-names>Horst</given-names>
          </name>
          <uri content-type="orcid">https://orcid.org/0000-0001-9407-3566</uri>
          <xref ref-type="aff" rid="A2">2</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Winterton</surname>
            <given-names>Shaun L.</given-names>
          </name>
          <xref ref-type="aff" rid="A3">3</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Aspöck</surname>
            <given-names>Ulrike</given-names>
          </name>
          <xref ref-type="aff" rid="A4">4</xref>
        </contrib>
      </contrib-group>
      <aff id="A1"><label>1</label>Department of Entomology, China Agricultural University, Beijing 100193, China
            </aff>
      <aff id="A2"><label>2</label>Institute of Specific Prophylaxis and Tropical Medicine, Medical Parasitology, Medical University of Vienna, Kinderspitalgasse 15, A-1090 Vienna, Austria
            </aff>
      <aff id="A3"><label>3</label>C
            </aff>
      <aff id="A4"><label>4</label>alifornia State Collection of Arthropods, California Department of Food &amp; Agriculture, 3294 Meadowview Rd. Sacramento, California, USA
            </aff>
      <aff id="A5"><label>5</label>Naturhistorisches Museum Wien, Zweite Zoologische Abteilung, Burgring 7, A-1010 Vienna, Austria
            </aff>
      <aff id="A6"><label>6</label>Department of Integrative Zoology, University of Vienna, Althanstraße 14, A-1090 Vienna, Austria
            </aff>
      <author-notes>
        <fn fn-type="corresp">
          <p>Corresponding author: Xingyue Liu (<email xlink:type="simple">xingyue_liu@yahoo.com</email>)</p>
          <!--PageBreak-->
        </fn>
        <fn fn-type="edited-by">
          <p>Academic editor: Susanne Randolf</p>
        </fn>
      </author-notes>
      <pub-date pub-type="collection">
        <year>2014</year>
      </pub-date>
      <pub-date pub-type="epub">
        <day>02</day>
        <month>12</month>
        <year>2014</year>
      </pub-date>
      <volume>61</volume>
      <issue>2</issue>
      <fpage>141</fpage>
      <lpage>153</lpage>
      <uri content-type="arpha" xlink:href="http://openbiodiv.net/FF957A43-EC6D-2256-FFEB-852648328D18">FF957A43-EC6D-2256-FFEB-852648328D18</uri>
      <uri content-type="zoobank" xlink:href="http://zoobank.org/2D0F3083-8FC9-4380-AC40-BF28519D72E1">2D0F3083-8FC9-4380-AC40-BF28519D72E1</uri>
      <uri content-type="zenodo_dep_id" xlink:href="https://zenodo.org/record/575722">575722</uri>
      <history>
        <date date-type="received">
          <day>22</day>
          <month>10</month>
          <year>2014</year>
        </date>
        <date date-type="accepted">
          <day>26</day>
          <month>11</month>
          <year>2014</year>
        </date>
      </history>
      <permissions>
        <copyright-statement>Wei Zhang, Xingyue Liu, Horst Aspöck, Shaun L. Winterton, Ulrike Aspöck</copyright-statement>
        <license license-type="creative-commons-attribution" xlink:href="http://creativecommons.org/licenses/by/4.0/" xlink:type="simple">
          <license-p>This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.</license-p>
        </license>
      </permissions>
      <self-uri content-type="zoobank" xlink:type="simple">http://zoobank.org/2D0F3083-8FC9-4380-AC40-BF28519D72E1</self-uri>
      <abstract>
        <label>Abstract</label>
        <p>Six species of the pleasing lacewing genus <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> Rambur, 1838, are recorded from several islands of East Asia, including Hainan, Taiwan, and Japan. Five species, i.e. <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">formosanus</tp:taxon-name-part></tp:taxon-name></italic> (Okamoto &amp; Kuwayama, 1920), <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">hikosanus</tp:taxon-name-part></tp:taxon-name></italic> Nakahara, 1955, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">japonicus</tp:taxon-name-part></tp:taxon-name></italic> McLachlan, 1883, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">pallidus</tp:taxon-name-part></tp:taxon-name></italic> Nakahara, 1955 and <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">taiwanensis</tp:taxon-name-part></tp:taxon-name></italic> Banks, 1937, are herein redescribed. <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">distinctus</tp:taxon-name-part></tp:taxon-name></italic> Nakahara, 1955, and <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">kanoi</tp:taxon-name-part></tp:taxon-name></italic> Nakahara, 1955, are synonymized with <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">taiwanensis</tp:taxon-name-part></tp:taxon-name></italic> Banks, 1937. <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">insularis</tp:taxon-name-part></tp:taxon-name></italic><bold>sp. n.</bold> from Hainan and Taiwan is described as new to science. A key to these insular species of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> is also provided.</p>
      </abstract>
      <kwd-group>
        <label>Key Words</label>
        <kwd>
          <tp:taxon-name>
            <tp:taxon-name-part taxon-name-part-type="family">Dilaridae</tp:taxon-name-part>
          </tp:taxon-name>
        </kwd>
        <kwd>
          <italic>
            <tp:taxon-name>
              <tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part>
            </tp:taxon-name>
          </italic>
        </kwd>
        <kwd>new species</kwd>
        <kwd>islands</kwd>
        <kwd>East Asia</kwd>
      </kwd-group>
    </article-meta>
  </front>
  <body>
    <sec sec-type="Introduction" id="sec1">
      <title>Introduction</title>
      <p>The <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="family">Dilaridae</tp:taxon-name-part></tp:taxon-name>, a small neuropteran family comprising about 80 valid species, are sorted in five genera of two subfamilies, i.e. <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="superfamily">Dilarinae</tp:taxon-name-part></tp:taxon-name> (only occur in the Old World) and <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="subfamily">Nallachiinae</tp:taxon-name-part></tp:taxon-name> (mainly occur in the New World, with two species recorded from Vietnam and southern Africa) (<xref ref-type="bibr" rid="B4">Engel 1999</xref>; <xref ref-type="bibr" rid="B7">Monserrat 1988</xref>, <xref ref-type="bibr" rid="B9">1989</xref>, <xref ref-type="bibr" rid="B10">2005</xref>; <xref ref-type="bibr" rid="B15">Oswald 1998</xref>, <xref ref-type="bibr" rid="B17">2013</xref>). The genus <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> Rambur, 1838 is the most species rich genus of <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="family">Dilaridae</tp:taxon-name-part></tp:taxon-name>, belonging to the subfamily <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="superfamily">Dilarinae</tp:taxon-name-part></tp:taxon-name>, with 52 described species (<xref ref-type="bibr" rid="B15">Oswald 1998</xref>; <xref ref-type="bibr" rid="B16">Oswald and Schiff 2001</xref>; <xref ref-type="bibr" rid="B18">Yang 1992</xref>, <xref ref-type="bibr" rid="B19">1999</xref>, <xref ref-type="bibr" rid="B20">2001</xref>; <xref ref-type="bibr" rid="B1">Aspöck et al. 2001</xref>; <xref ref-type="bibr" rid="B21">Zhang et al. 2014a</xref>). Majority of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> species occur in the Eurasia continent, while a few species are distributed in some islands of Asia and Europe. For instance, Sri Lanka harbours an endemic species, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">nietneri</tp:taxon-name-part></tp:taxon-name></italic> Hagen, 1858, and two species, i.e. <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">grandis</tp:taxon-name-part></tp:taxon-name></italic> (Banks, 1931) and <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">macleodi</tp:taxon-name-part></tp:taxon-name></italic> Oswald &amp; Schiff, 2001 only occur in Borneo. In East Asia, there are many islands along the coast, including Hainan (second largest island of China), Taiwan (the largest island of China), Ryukyus, and Japanese archipelago. Hitherto, seven species of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> have been recorded from some of these islands (<xref ref-type="bibr" rid="B6">McLachlan 1883</xref>; <xref ref-type="bibr" rid="B14">Okamoto and Kuwayama 1920</xref>; <xref ref-type="bibr" rid="B3">Banks 1937</xref>; <xref ref-type="bibr" rid="B11">Nakahara 1955a</xref>, <xref ref-type="bibr" rid="B12">b</xref>). Among these species, two of them are distributed in Japan, while, remarkably, five species were described and endemic to Taiwan. However, all these seven <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> species were poorly described and lack a clear, modern redescription and illustrations. In this paper, we revise the insular species of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> from East Asia. Six valid species are herein recorded and described from Hainan, Taiwan, and Japan, with one new species. A key to the insular species of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> from East Asia is also given.</p>
    </sec>
    <sec sec-type="materials|methods" id="sec2">
      <title>Material and methods</title>
      <p>Specimens for the present study are deposited in the Natural History Museum (BMNH), London, U.K.; the Entomological Museum of China Agricultural University (CAU), Beijing, China; the Museum of Comparative Zoology (MCZ), Cambridge, U.S.A; Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung an der Humboldt-Universität zu Berlin (MNB), Berlin, Germany; National Museum of Natural History (NMNH), Smithsonian Institutions, Washington, D.C., U.S.A.; the National Museum of Nature and Science (NSMT), Tsukuba, Japan; the National Institute for Agro-Environmental Sciences (NIAES), Tsukuba, Japan; and the Entomological Collection of Hokkaido University Museum (SEHU), Sapporo, Japan. Genitalic preparations were made by clearing the apex of the abdomen in a cold, saturated KOH solution for 3-4 h. After rinsing the KOH with acetic acid and water, the apex of the abdomen was transferred to glycerin for further dissection and examination. Habitus photos of adults were taken by using a Nikon D90 digital camera with Nikon MICRO NIKKOR 105 mm lens, and the genitalic figures were made by hand drawing under a Motic SMZ168 stereo microscope. The terminology of the genitalia generally follows U. Aspöck and H. Aspöck (<xref ref-type="bibr" rid="B2">2008</xref>).</p>
    </sec>
    <sec sec-type="Taxonomy" id="sec3">
      <title>Taxonomy</title>
      <tp:taxon-treatment>
        <tp:treatment-meta>
          <kwd-group>
            <label>Taxon classification</label>
            <kwd>
              <named-content content-type="kingdom" xlink:type="simple">Animalia</named-content>
            </kwd>
          </kwd-group>
        </tp:treatment-meta>
        <tp:nomenclature>
          <label>Genus</label>
          <tp:taxon-name><object-id content-type="arpha">DF4E03C5-9AB7-5710-83DD-9F89A17D994E</object-id>
            <tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part>
          </tp:taxon-name>
          <tp:taxon-authority>Rambur</tp:taxon-authority>
          <tp:nomenclature-citation-list>
            <tp:nomenclature-citation>
              <tp:taxon-name>
                <tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part>
              </tp:taxon-name>
              <comment> Rambur, 1838: 9. Type species: <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">nevadensis</tp:taxon-name-part></tp:taxon-name></italic> Rambur, 1838 (monotypy).</comment>
            </tp:nomenclature-citation>
            <tp:nomenclature-citation>
              <tp:taxon-name>
                <tp:taxon-name-part taxon-name-part-type="genus">Cladocera</tp:taxon-name-part>
              </tp:taxon-name>
              <comment> Hagen, 1860: 56. Nomen nudum.</comment>
            </tp:nomenclature-citation>
            <tp:nomenclature-citation>
              <tp:taxon-name>
                <tp:taxon-name-part taxon-name-part-type="genus">Lidar</tp:taxon-name-part>
              </tp:taxon-name>
              <comment> Navás, 1909: 153. Type species: <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">meridionalis</tp:taxon-name-part></tp:taxon-name></italic> Hagen, 1866: 295, original designation.</comment>
            </tp:nomenclature-citation>
            <tp:nomenclature-citation>
              <tp:taxon-name>
                <tp:taxon-name-part taxon-name-part-type="genus">Fuentenus</tp:taxon-name-part>
              </tp:taxon-name>
              <comment> Navás, 1909: 154. Type species: <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">campestris</tp:taxon-name-part></tp:taxon-name></italic> Navás, 1903: 380, original designation.</comment>
            </tp:nomenclature-citation>
            <tp:nomenclature-citation>
              <tp:taxon-name>
                <tp:taxon-name-part taxon-name-part-type="genus">Nepal</tp:taxon-name-part>
              </tp:taxon-name>
              <comment> Navás, 1909: 661. Type species: <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Nepal</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">harmandi</tp:taxon-name-part></tp:taxon-name></italic> Navás, 1909: 661, original designation.</comment>
            </tp:nomenclature-citation>
            <tp:nomenclature-citation>
              <tp:taxon-name>
                <tp:taxon-name-part taxon-name-part-type="genus">Rexavius</tp:taxon-name-part>
              </tp:taxon-name>
              <comment> Navás, 1909: 664. Type species: <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">nietneri</tp:taxon-name-part></tp:taxon-name></italic> Hagen, 1858: 482, subsequent designation by Navás, 1914: 10.</comment>
            </tp:nomenclature-citation>
            <tp:nomenclature-citation>
              <tp:taxon-name>
                <tp:taxon-name-part taxon-name-part-type="genus">Didar</tp:taxon-name-part>
              </tp:taxon-name>
              <comment> Navás, 1913: 6. An incorrect subsequent spelling of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic>.</comment>
            </tp:nomenclature-citation>
            <tp:nomenclature-citation>
              <tp:taxon-name>
                <tp:taxon-name-part taxon-name-part-type="genus">Lider</tp:taxon-name-part>
              </tp:taxon-name>
              <comment> Kuwayama, 1962: 376. An incorrect subsequent spelling of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Lidar</tp:taxon-name-part></tp:taxon-name></italic>.</comment>
            </tp:nomenclature-citation>
          </tp:nomenclature-citation-list>
        </tp:nomenclature>
        <tp:treatment-sec sec-type="Diagnosis">
          <title>Diagnosis.</title>
          <p>Male antennae pectinate, except 2 proximal and more than 3 distal simple antennomeres, branch of 1st flagellomere short, with only one dentate process. Wings broad, generally with numerous dark markings; costal area broad with crossveins simple except several ones forked; subcostal area obviously narrower than costal area, having several crossveins; more than 5 crossveins between R and Rs, MA fused with R at wing base, proximally with no crossvein connecting to MP, and arising from R prior to separation between R and Rs; MP with 2 main branches; nygmata present; trichosors present along wing margin. Male ninth tergite in dorsal view with a truncate or arcuate anterior incision and a deeply V- or U-shaped posterior incision, leaving a pair of broad hemitergites, which are obtuse distally and densely haired. Male ninth sternite generally much shorter than ninth tergite. Male ectoproct highly specialized, largely covered by ninth tergite, without callus cerci and any macrosetae. Male gonocoxite complexes 9, 10 and 11 comprising two pairs of sclerites (i.e. ninth and tenth gonocoxites) and a transverse sclerite (i.e. gonarcus = eleventh gonocixites); gonarcus laterally connecting to bases of ninth gonocoxites. Hypandrium internum generally trapezoidal, with lateral margins slightly arcuate.</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Distribution">
          <title>Distribution.</title>
          <p>This genus ranges from northern Africa, through Europe, to Asia, and is recorded in the following countries: Afghanistan, Algeria, Andorra, Bulgaria, China, France, Greece, India, Iran, Italy, Japan, Korea, Kyrgyzstan, Lebanon, Malaysia, Nepal, Pakistan, Portugal, Russia, Spain, Sri Lanka, Tajikistan, Thailand, Turkey, Turkmenistan, Vietnam all countries of former Yugoslavia.</p>
          <sec sec-type="Key to males of insular Dilar species from East Asia" id="sec4">
            <title>Key to males of insular <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> species from East Asia</title>
            <table-wrap content-type="key" position="anchor" orientation="portrait">
              <table rules="all">
                <tbody>
                  <tr>
                    <td rowspan="1" colspan="1">1</td>
                    <td rowspan="1" colspan="1">Forewings nearly hyaline, with no obviously markings</td>
                    <td rowspan="1" colspan="1"><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">pallidus</tp:taxon-name-part></tp:taxon-name></italic> Nakahara</td>
                  </tr>
                  <tr>
                    <td rowspan="1" colspan="1">–</td>
                    <td rowspan="1" colspan="1">Forewings yellowish brown, with distinct brown markings</td>
                    <td rowspan="1" colspan="1">2</td>
                  </tr>
                  <tr>
                    <td rowspan="1" colspan="1">2</td>
                    <td rowspan="1" colspan="1">Male (Fig. <xref ref-type="fig" rid="F7">24</xref>, <xref ref-type="fig" rid="F7">26</xref>) with ninth tergite in dorsal view with posteromedial portion protruding into a projection</td>
                    <td rowspan="1" colspan="1">
                      <italic>
                        <tp:taxon-name>
                          <tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part>
                          <tp:taxon-name-part taxon-name-part-type="species">insularis</tp:taxon-name-part>
                        </tp:taxon-name>
                      </italic>
                      <bold>sp. n.</bold>
                    </td>
                  </tr>
                  <tr>
                    <td rowspan="1" colspan="1">–</td>
                    <td rowspan="1" colspan="1">Male with ninth tergite in dorsal view without projection posteromedially</td>
                    <td rowspan="1" colspan="1">3</td>
                  </tr>
                  <tr>
                    <td rowspan="1" colspan="1">3</td>
                    <td rowspan="1" colspan="1">Male (Fig. <xref ref-type="fig" rid="F4">17</xref>) with ectoproct in dorsal view with a strong sclerotized projection at tip and with gonarcus beam-shaped, which is bifid on both ends</td>
                    <td rowspan="1" colspan="1"><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">taiwanensis</tp:taxon-name-part></tp:taxon-name></italic> Banks</td>
                  </tr>
                  <tr>
                    <td rowspan="1" colspan="1">–</td>
                    <td rowspan="1" colspan="1">Male with gonarcus beam-shaped, which is not bifid on both ends</td>
                    <td rowspan="1" colspan="1">4</td>
                  </tr>
                  <tr>
                    <td rowspan="1" colspan="1">4</td>
                    <td rowspan="1" colspan="1">Male (Fig. <xref ref-type="fig" rid="F3">13</xref>) with ectoproct in dorsal view with a pair of parallel prolongations which are truncatus and serrate at tip</td>
                    <td rowspan="1" colspan="1"><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">hikosanus</tp:taxon-name-part></tp:taxon-name></italic> Nakahara</td>
                  </tr>
                  <tr>
                    <td rowspan="1" colspan="1">–</td>
                    <td rowspan="1" colspan="1">Male (Fig. <xref ref-type="fig" rid="F2">7</xref>) with ectoproct in dorsal view with three projections at tip which is sparsely haired</td>
                    <td rowspan="1" colspan="1"><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">japonicus</tp:taxon-name-part></tp:taxon-name></italic> McLachlan</td>
                  </tr>
                  <!--PageBreak-->
                </tbody>
              </table>
            </table-wrap>
          </sec>
        </tp:treatment-sec>
      </tp:taxon-treatment>
      <tp:taxon-treatment>
        <tp:treatment-meta>
          <kwd-group>
            <label>Taxon classification</label>
            <kwd>
              <named-content content-type="kingdom" xlink:type="simple">Animalia</named-content>
            </kwd>
          </kwd-group>
        </tp:treatment-meta>
        <tp:nomenclature>
          <tp:taxon-name><object-id content-type="arpha">218F3AE1-4BD8-5B09-A13B-918351CF019A</object-id>
            <tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part>
            <tp:taxon-name-part taxon-name-part-type="species">japonicus</tp:taxon-name-part>
          </tp:taxon-name>
          <tp:taxon-authority>McLachlan, 1883</tp:taxon-authority>
          <xref ref-type="fig" rid="F1">Figs 1</xref>
          <xref ref-type="fig" rid="F2">, 7–12</xref>
          <tp:nomenclature-citation-list>
            <tp:nomenclature-citation>
              <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">japonicus</tp:taxon-name-part></tp:taxon-name>
              <comment> McLachlan, 1883: 220. Type locality: Japan (Fukushima).</comment>
            </tp:nomenclature-citation>
            <tp:nomenclature-citation>
              <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">nohirae</tp:taxon-name-part></tp:taxon-name>
              <comment> Nakahara, 1914: 297. Type locality: Japan (Nara).</comment>
            </tp:nomenclature-citation>
            <tp:nomenclature-citation>
              <tp:taxon-name>
                <tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar"/>
              </tp:taxon-name>
              <comment>Dilar japonicus var. gracilis Kuwayama, 1921: 71. Type locality: Japan (Fukushima).</comment>
            </tp:nomenclature-citation>
          </tp:nomenclature-citation-list>
        </tp:nomenclature>
        <tp:treatment-sec sec-type="Diagnosis">
          <title>Diagnosis.</title>
          <p>This species is characterized by the forewings with numerous pale brown markings, and the male ectoproct in dorsal view with three projections at tip which is sparsely haired.</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Description">
          <title>Description.</title>
          <p>Male. Body length 4.3-5.3 mm; forewing length 10.0-11.4 mm, hindwing length 8.8-9.4 mm.</p>
          <p>Head pale yellowish brown, with pale yellow setose tubercles. Compound eyes blackish brown. Antenna with ca. 29 segments, pale yellowish brown, pedicel with brown annular stripes, flagellum unipectinate on most flagellomeres, medial branches much longer than those on both ends, longest branch nearly 3.0 times as long as relevant flagellomere, but branch of 1st flagellomere short and dentate, distal seven flagellomeres simple.</p>
          <p>Prothorax pale yellow, pronotum pale yellowish brown, with anterior margin and posterolateral corners pale yellow, medially with a pair of ovoid markings; mesothorax pale yellowish brown, mesonotum dark brown on anterior and lateral margins; metanotum pale yellowish brown, slightly darker on lateral margins. Legs pale yellowish brown, femora blackish brown at tip. Wings hyaline, slightly smoky brown, with numerous brown spots. Forewing ~2.1 times as long as wide, densely spotted, with markings on proximal half much darker, arranging as transversely arcuate pattern, a broad immaculate area present on either side of median nygma; two nygmata present on proximal and median portion of forewing, median one much larger than proximal one. Hindwing ~2.0 times as long as wide, slightly paler than forewing, with similar marking pattern; one nygma present at middle. Veins pale brown, crossveins much paler than longitudinal veins. Forewing with trichosors present along wing margin between R and CuA; Rs with three main branches; MP with two main branches; two gradate crossveins present at middle. Hindwing with trichosors present along wing margin between R and CuA; Rs with three main branches.</p>
          <p>Abdomen pale yellow, pregenital segments dorsally yellowish brown. Ninth tergite in dorsal view with an arcuate anterior incision and a deeply V-shaped posterior incision, leaving a narrow median portion and a pair of subtriangular hemitergites, which are obtuse distally and densely haired; in lateral view broad, with straight ventral margin and arcuate posterior margin. Ninth sternite obviously shorter than ninth tergite, truncately posteriad. Ectoproct in dorsal view with three projections at tip which is sparsely haired, posteroventrally with a pair of subsemicircular and flattened projections, a pair of bifid unguiform projections and a pair of short, feebly sclerotized, digitiform projections. Ninth gonocoxite with anterior half broadly spoon-shaped and with posterior half slenderly elongate and curved posteroventrally; tenth gonocoxite slenderly elongate, much longer ninth gonocoxite, with incurved base and spinous tip, medially with a lobe connecting to ninth gonocoxite; gonarcus beam-shaped, slightly expanded on both ends, laterally connecting to base of ninth gonocoxites. Hypandrium internum nearly trapezoidal, with lateral margins slightly arcuate.</p>
          <p>Female. Body length 6.5 mm; forewing length 14.0 mm, hindwing length 12.7 mm.</p>
          <p>Seventh sternite (Fig. <xref ref-type="fig" rid="F2">11</xref>) in lateral view subtrapezoidal, in ventral view nearly rectangular and slightly convex on posterior margin. Eighth abdominal segment ventrally without subgenitale. Bursa copulatrix (Fig. <xref ref-type="fig" rid="F2">11</xref>) with colleterial gland tubular and elongate, sinuate medially; basal part of bursa copulatrix sac-like in lateral view and nearly ovoid in ventral view, posteriorly slightly sclerotized on both sides; bursal accessory gland not observed. Ectoproct rather small, ovoid.</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Materials examined">
          <title>Materials examined.</title>
          <p>Holotype ♂, “Japan, Fukushima [<named-content content-type="dwc:verbatimCoordinates">37°44’N, 140°28’E</named-content>]/28.7.[18]81/Type/<italic>japonicus</italic> McL[achlan]” (BMNH). JAPAN: 1f#, Aomori, Tsuta-onsen [<named-content content-type="dwc:verbatimCoordinates">40°13’N, 140°30’E</named-content>], 6.VIII.1953, K. Sadanaga (NIAES); 1♀, Gunma, Kuni-mura [<named-content content-type="dwc:verbatimCoordinates">36°38’N, 138°38’E</named-content>], 10.VIII.1992, K. Konishi (NIAES); 1♂, Gunma [<named-content content-type="dwc:verbatimCoordinates">36°23’N, 139°03’E</named-content>], 3.VIII.1960, K. Fujimoto (NIAES); 1♂, Shikoku, Mt. Sara [<named-content content-type="dwc:verbatimCoordinates">33°45’N, 133°30’E</named-content>], 27.VII.1955, Toshiro Yano (NIAES); 1♂, Yamanashi, Shojiko [<named-content content-type="dwc:verbatimCoordinates">35°29’N, 138°36’E</named-content>], 23.VII.1969, P. H. Hasegawa (NIAES); 1f#, Fukushima, Yokomuki Onsen [<named-content content-type="dwc:verbatimCoordinates">37°53’N, 140°33’E</named-content>], 26.VII.1964, H. Kobayashi (NIAES).</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Distribution">
          <title>Distribution.</title>
          <p>Japan (Honshu, Kyushu, Shikoku).</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Remarks">
          <title>Remarks.</title>
          <p>This species appears to be very closely related to <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">hikosanus</tp:taxon-name-part></tp:taxon-name></italic> Nakahara in having similar wing marking patterns and male genitalia. The only difference between these two Japanese species refers to the male ectoproct. In <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">japonicus</tp:taxon-name-part></tp:taxon-name></italic>, the male ectoproct distally bears three projections which are sparsely haired. However, in <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">hikosanus</tp:taxon-name-part></tp:taxon-name></italic> the ectoproct has a pair of parallel projections which are truncate and serrate at tip. Primarily we even suspected <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">hikosanus</tp:taxon-name-part></tp:taxon-name></italic> to be conspecific with <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">japonicus</tp:taxon-name-part></tp:taxon-name></italic> due to the remarkable similarity of the external morphological and genitalic characters. Nevertheless, we examined several males of both species, and the difference of the male ectoproct mentioned above is stable. Therefore, we consider <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">hikosanus</tp:taxon-name-part></tp:taxon-name></italic> and <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">japonicus</tp:taxon-name-part></tp:taxon-name></italic> to be different species. <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">japonicus</tp:taxon-name-part></tp:taxon-name></italic> is a widely distributed species in Honshu, Kyushu and Shikoku, while <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">hikosanus</tp:taxon-name-part></tp:taxon-name></italic> seems to be endemic to Kyushu, and more exactly to the Mt. Hikosan.</p>
          <fig id="F1" position="float" orientation="portrait">
            <object-id content-type="doi">10.3897/dez.61.8793.figures1-6</object-id>
            <object-id content-type="zenodo_dep_id">959859</object-id>
            <object-id content-type="arpha">C8D67BD1-CC13-52B0-81E5-369554F53B23</object-id>
            <label>Figures 1–6.</label>
            <caption>
              <p>Adults of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> spp. <bold>1.</bold><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">japonicus</tp:taxon-name-part></tp:taxon-name></italic> McLachlan, holotype male; <bold>2.</bold><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">hikosanus</tp:taxon-name-part></tp:taxon-name></italic> Nakahara, male; <bold>3.</bold><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">taiwanensis</tp:taxon-name-part></tp:taxon-name></italic> Banks, holotype male; <bold>4.</bold><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">formosanus</tp:taxon-name-part></tp:taxon-name></italic> (Okamoto &amp; Kuwayama), holotype female, fore- and hindwing; <bold>5.</bold><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">pallidus</tp:taxon-name-part></tp:taxon-name></italic> Nakahara, holotype male; <bold>6.</bold><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">insularis</tp:taxon-name-part></tp:taxon-name></italic> sp. n., holotype male. Scale bars: 1.0 mm.</p>
            </caption>
            <graphic xlink:href="dez-61-141-g001.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_34977.jpg">
              <uri content-type="original_file">https://binary.pensoft.net/fig/34977</uri>
            </graphic>
          </fig>
          <fig id="F2" position="float" orientation="portrait">
            <object-id content-type="doi">10.3897/dez.61.8793.figures7-12</object-id>
            <object-id content-type="zenodo_dep_id">959861</object-id>
            <object-id content-type="arpha">795CBBB6-C083-5320-B974-3EED6DC88E0A</object-id>
            <label>Figures 7–12.</label>
            <caption>
              <p><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">japonicus</tp:taxon-name-part></tp:taxon-name></italic> McLachlan. <bold>7.</bold> Male genitalia, dorsal view; <bold>8.</bold> Male genitalia, ventral view; <bold>9.</bold> Male genitalia, lateral view; <bold>10.</bold> Male ectoproct, caudal view; <bold>11.</bold> Female genitalia, lateral view; <bold>12.</bold> Female genitalia, ventral view. bc: bursa copulatrix; cg: colleterial gland; e: ectoproct; g: gonarcus = eleventh gonocoxites; gx9: ninth gonocoxite; gx10: tenth gonocoxite; hi: hypandrium internum; S7-9: sternite 7-9; T7-9: tergite 7-9. Scale bars: 0.5 mm.</p>
            </caption>
            <graphic xlink:href="dez-61-141-g002.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_34978.jpg">
              <uri content-type="original_file">https://binary.pensoft.net/fig/34978</uri>
            </graphic>
          </fig>
        </tp:treatment-sec>
      </tp:taxon-treatment>
      <tp:taxon-treatment>
        <tp:treatment-meta>
          <kwd-group>
            <label>Taxon classification</label>
            <kwd>
              <named-content content-type="kingdom" xlink:type="simple">Animalia</named-content>
            </kwd>
          </kwd-group>
        </tp:treatment-meta>
        <tp:nomenclature>
          <tp:taxon-name><object-id content-type="arpha">CD5E527A-BF31-59DC-8A0D-C9C945418FD9</object-id>
            <tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part>
            <tp:taxon-name-part taxon-name-part-type="species">hikosanus</tp:taxon-name-part>
          </tp:taxon-name>
          <tp:taxon-authority>Nakahara, 1955</tp:taxon-authority>
          <xref ref-type="fig" rid="F1">Figs 2</xref>
          <xref ref-type="fig" rid="F3">, 13</xref>
          <xref ref-type="fig" rid="F3">–16</xref>
          <tp:nomenclature-citation-list>
            <tp:nomenclature-citation>
              <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">hikosanus</tp:taxon-name-part></tp:taxon-name>
              <comment> Nakahara, 1955b: 137. Type locality: Japan (Fukuoka).</comment>
            </tp:nomenclature-citation>
          </tp:nomenclature-citation-list>
        </tp:nomenclature>
        <tp:treatment-sec sec-type="Diagnosis">
          <title>Diagnosis.</title>
          <p>This species is characterized by the forewings with numerous brown stripes, and the male ectoproct in dorsal view with a pair of parallel prolongations which are truncate and serrate at tip.</p>
          <p><bold>Description.</bold> Male. Body length 4.7 mm; forewing length 8.3 mm, hindwing length 6.8 mm.</p>
          <p>Head yellowish brown, with pale yellow setose tubercles. Compound eyes blackish brown. Antenna with ca. 29 segments, pale yellowish brown, pedicel with brown annular stripes, flagellum unipectinate on most flagellomeres, medial branches much longer than those on both ends, longest branch nearly 3.0 times as long as relevant flagellomere, but branch of 1st flagellomere short and dentate, distal eight flagellomeres simple.</p>
          <p>Prothorax pale yellowish brown, pronotum yellowish brown, with anterior margin and posterolateral corners pale yellow, medially with a pair of ovoid markings; mesothorax pale yellowish brown, mesonotum dark brown on anterior and lateral margins; metanotum pale yellowish brown, slightly darker on lateral margins. Legs pale yellowish brown, femora blackish brown at tip. Wings hyaline, slightly yellowish brown, with numerous brown spots. Forewing ~2.2 times as long as wide, densely spotted, with markings on proximal half much darker, most markings connected with each other, generally arranging into several arcuate transverse stripes, an immaculate area present distal to median nygma; two nygmata present on proximal and median portion of forewing, median one much larger than proximal one. Hindwing ~2.1 times as long as wide, slightly paler than forewing, with similar marking pattern; one nygma present at middle. Veins pale brown. Forewing with trichosors present along wing margin between R and CuP; Rs with four main branches; MP with two main branches; two gradate crossveins present at middle. Hindwing with trichosors present along wing margin between R and CuA; Rs with three main branches.</p>
          <p>Abdomen pale yellowish brown, pregenital segments dorsally brown. Ninth tergite in dorsal view with an arcuate anterior incision and a nearly V-shaped posterior incision, leaving a narrow median portion and a pair of subtriangular hemitergites, which are obtuse distally and densely haired; in lateral view broad, with straight ventral margin and arcuate posterior margin. Ninth sternite obviously shorter than ninth tergite, arcuately convex posteriad. Ectoproct in dorsal view with a pair of parallel prolongations which are truncate and serrate at tip, posteroventrally with a pair of subsemicircular and flattened projections, a pair of bifid unguiform projections and a pair of short, feebly sclerotized, digitiform projections. Ninth gonocoxite with anterior half broadly spoon-shaped and with posterior half slenderly elongate and curved posteroventrally; tenth gonocoxite slenderly elongate, much longer ninth gonocoxite, with incurved base and spinous tip which is curved outward, medially with a lobe connecting to ninth gonocoxite; gonarcus beam-shaped, laterally connecting to base of ninth gonocoxites. Hypandrium internum nearly trapezoidal, with lateral margins slightly arcuate.</p>
          <p>Female. Unknown.</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Materials examined">
          <title>Materials examined.</title>
          <p>Holotype ♂, “HIKOSAN [<named-content content-type="dwc:verbatimCoordinates">33°01’N, 131°01’E</named-content>], Kyushu, Aug. 3, 1934, S. Hiroyama/<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">hikosanus</tp:taxon-name-part></tp:taxon-name></italic> n.sp. (Type) W. NAKAHARA/Waro Nakahara Collection II/<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">hikosanus</tp:taxon-name-part></tp:taxon-name></italic> n.sp. W. NAKAHARA (Holotype)/NSMT-I-Nr No. 4302” (NSMT). 2♂, JAPAN: Kyushu, Hikosan, Buzen, 19.VII.1963, H. Kuroko (NIAES).</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Distribution">
          <title>Distribution.</title>
          <p>Japan (Kyushu).</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Remarks">
          <title>Remarks.</title>
          <p>See Remarks under <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">japonicus</tp:taxon-name-part></tp:taxon-name></italic> McLachlan, 1883.</p>
          <fig id="F3" position="float" orientation="portrait">
            <object-id content-type="doi">10.3897/dez.61.8793.figures13-16</object-id>
            <object-id content-type="zenodo_dep_id">959863</object-id>
            <object-id content-type="arpha">4D40A1D1-41D2-5CBF-AFFB-878A5F2BAACF</object-id>
            <label>Figures 13–16.</label>
            <caption>
              <p><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">hikosanus</tp:taxon-name-part></tp:taxon-name></italic> Nakahara. <bold>13.</bold> Male genitalia, dorsal view; <bold>14.</bold> Male genitalia, ventral view; <bold>15.</bold> Male genitalia, lateral view; <bold>16.</bold> Male ectoproct, caudal view. Scale bars: 0.5 mm.</p>
            </caption>
            <graphic xlink:href="dez-61-141-g003.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_34979.jpg">
              <uri content-type="original_file">https://binary.pensoft.net/fig/34979</uri>
            </graphic>
          </fig>
          <!--PageBreak-->
        </tp:treatment-sec>
      </tp:taxon-treatment>
      <tp:taxon-treatment>
        <tp:treatment-meta>
          <kwd-group>
            <label>Taxon classification</label>
            <kwd>
              <named-content content-type="kingdom" xlink:type="simple">Animalia</named-content>
            </kwd>
          </kwd-group>
        </tp:treatment-meta>
        <tp:nomenclature>
          <tp:taxon-name><object-id content-type="arpha">2711BFBD-FA3D-5868-A45C-9C716D93A8E1</object-id>
            <tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part>
            <tp:taxon-name-part taxon-name-part-type="species">taiwanensis</tp:taxon-name-part>
          </tp:taxon-name>
          <tp:taxon-authority>Banks, 1937</tp:taxon-authority>
          <xref ref-type="fig" rid="F1">Figs 3</xref>
          <xref ref-type="fig" rid="F4">, 17</xref>
          <xref ref-type="fig" rid="F4">–20</xref>
          <!--PageBreak-->
          <!--PageBreak-->
          <!--PageBreak-->
          <tp:nomenclature-citation-list>
            <tp:nomenclature-citation>
              <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">taiwanensis</tp:taxon-name-part></tp:taxon-name>
              <comment> Banks, 1937: 276. Type locality: China (Taiwan: Jiayi).</comment>
            </tp:nomenclature-citation>
            <tp:nomenclature-citation>
              <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">distinctus</tp:taxon-name-part></tp:taxon-name>
              <comment> Nakahara, 1955b: 139. Type locality: China (Taiwan: Tattaka). <bold>syn. n.</bold></comment>
            </tp:nomenclature-citation>
            <tp:nomenclature-citation>
              <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Nepal</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">kanoi</tp:taxon-name-part></tp:taxon-name>
              <comment> Nakahara, 1955a: 6. Type locality: China (Taiwan: Tattaka). <bold>syn. n.</bold></comment>
            </tp:nomenclature-citation>
          </tp:nomenclature-citation-list>
        </tp:nomenclature>
        <tp:treatment-sec sec-type="Diagnosis">
          <title>Diagnosis.</title>
          <p>This species is characterized by the male ninth gonocoxite with anterior half broadly spoon-shaped and with posterior half slenderly elongate and curved posteroventrally, and by the slenderly elongate tenth gonocoxite which has a lobe connecting to ninth gonocoxite medially.</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Description">
          <title>Description.</title>
          <p>Male. Body length 3.4–5.5 mm; forewing length 6.4–9.0 mm, hindwing length 5.5–7.6 mm.</p>
          <p>Head yellowish brown, with pale yellow setose tubercles. Compound eyes blackish brown. Antenna with ca. 29 segments, pale yellowish brown, pedicel with brown annular stripes, flagellum unipectinate on most flagellomeres, medial branches much longer than those on both ends, longest branch nearly 3.0 times as long as relevant flagellomere, but branch of 1st flagellomere short and dentate, distal eight flagellomeres simple.</p>
          <p>Prothorax pale yellowish brown, pronotum yellowish brown, with anterior margin and posterolateral corners pale yellow, medially with a pair of ovoid markings; mesothorax pale yellowish brown, mesonotum dark brown on anterior and lateral margins; metanotum pale yellowish brown, slight darker on lateral margins. Legs pale yellowish brown, femora blackish brown at tip. Wings hyaline, slightly yellowish brown, with numerous dark spots. Forewing ~2.2 times as long as wide, densely spotted, with markings on proximal half much darker, most markings fused with each other, generally arranging into several arcuate transverse stripes, an immaculate area present distal to median nygma; two nygmata present on proximal and median portion of forewing, median one much larger than proximal one. Hindwing ~2.1 times as long as wide, slightly paler than forewing, with similar marking pattern; one nygma present at middle. Veins pale brown. Forewing with trichosors present along wing margin between R and CuP; Rs with four main branches; MP with two main branches; two gradate crossveins present at middle. Hindwing with trichosors present along wing margin between R and CuA; Rs with three main branches.</p>
          <p>Abdomen pale yellowish brown, pregenital segments dorsally brown. Ninth tergite in dorsal view with an arcuate anterior incision and a nearly V-shaped posterior incision, leaving a narrow median portion and a pair of subtriangular hemitergites, which are obtuse distally and densely haired; in lateral view broad, with straight ventral margin and arcuate posterior margin. Ninth sternite obviously shorter than ninth tergite, arcuately convex posteriad. Ectoproct in dorsal view with a strong sclerotized projection medially, which is bifid at tip, posteroventrally with a pair of short and flattened projections, a pair of bifid unguiform projections and a pair of short, feebly sclerotized, digitiform projections. Ninth gonocoxite strongly inflated and spoon-shaped on anterior half, with posterior half slenderly elongate and curved posteroventrally, which is with a slenderly short projection extended subterminally; tenth gonocoxite slenderly elongate, much longer than ninth gonocoxite, with incurved base and spinous tip which is curved outward, medially with a lobe connecting to ninth gonocoxite; gonarcus beam-shaped, bifid on both ends, laterally connecting to base of ninth gonocoxites. Hypandrium internum nearly trapezoidal, with lateral margins slightly arcuate.</p>
          <p>Female. Forewing length 12.5 mm, hindwing length 11.5 mm.</p>
          <p>Abdomen yellow, ovipositor yellowish brown, seventh sternite ruptured during maceration, eighth abdominal segment without subgenitale. Colleterial gland apparently also ruptured, not visible. Bursa copulatrix sac-like in lateral view, with an elongated bowl-like sclerite and a paired sclerotized domed structure distally behind it; bursal accessory gland not observed. Ectoproct rather small, ovoid.</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Materials examined">
          <title>Materials examined.</title>
          <p>Holotype ♂, “Formosa, Arizan [= Mt. Alishan, Taiwan, <named-content content-type="dwc:verbatimCoordinates">23°31’N, 120°48’E</named-content>], June 4, Gressitt, Type, M.C.Z. No. 20229” (MCZ). CHINA: 2♂, Taiwan, Taoyuan, Mt. Tamanshan [<named-content content-type="dwc:verbatimCoordinates">24°42’N, 121°27’E</named-content>], 1600 m, 11.VI.2013, Feiyang Liang (CAU); 1♂, Taiwan, Taipei, Mt. Yangmingshan, Datun Park [<named-content content-type="dwc:verbatimCoordinates">25°09’N, 121°32’E</named-content>], 900 m, 17.VI.2012, Lihua Wang (CAU); 2♂, Taiwan, Taipei, Mt. Yangmingshan, Datun Park, 900 m, 17.VI.2012, Luxi Liu (CAU); 5♂1f#, Taiwan, Nantou, Tayuling [<named-content content-type="dwc:verbatimCoordinates">23°55’N, 120°40’E</named-content>], 2500 m, 8-18.VI.1998, D. R. Davis (NMNH); 1♂, “Formosa, Taihorin [Taiwan: Chiayi County, Dalin (= Taihorin), <named-content content-type="dwc:verbatimCoordinates">23°35’N, 120°28’E</named-content>], VI.10. [= June 1910], H. Sauter” (MNB); 2♂ [Holotype and a paratype of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">distinctus</tp:taxon-name-part></tp:taxon-name></italic>], “Formosa, T. Kano/<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">distinctus</tp:taxon-name-part></tp:taxon-name></italic> n.sp. (Type) W. NAKAHARA/Waro Nakahara Collection II” (NSMT); 1♂ [Holotype of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">kanoi</tp:taxon-name-part></tp:taxon-name></italic>], “Formosa, T. Kano/<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">kanoi</tp:taxon-name-part></tp:taxon-name></italic> n.sp. (Type) W. NAKAHARA/Waro Nakahara Collection II/NSMT-I-Nr No. 4340” (NSMT); 1♂, CHINA: Taiwan, 29.VII.1928, T. Kano (NSMT); JAPAN: 1♂, Iriomote Island, Upper Nakara River [<named-content content-type="dwc:verbatimCoordinates">24°20’N, 123°49’E</named-content>], 12.III.1964, Y. Miyatake (NSMT).</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Distribution">
          <title>Distribution.</title>
          <p>China (Taiwan), Japan (Iriomote Island).</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Remarks">
          <title>Remarks.</title>
          <p>This species was originally described by <xref ref-type="bibr" rid="B3">Banks (1937)</xref> based on external morphology only. After our examination of the holotype of this species, we found that this species is identical with <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">distinctus</tp:taxon-name-part></tp:taxon-name></italic> Nakahara and <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">kanoi</tp:taxon-name-part></tp:taxon-name></italic> Nakahara based on the comparison of its male genitalia with the original genitalic illustrations of the latter two species. Therefore, we treat <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">distinctus</tp:taxon-name-part></tp:taxon-name></italic> and <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">kanoi</tp:taxon-name-part></tp:taxon-name></italic> as junior synonyms of <italic>D. taiwanensis.</italic> Nevertheless, it should be noted that the collecting labels of the primary types of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">distinctus</tp:taxon-name-part></tp:taxon-name></italic> and <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">kanoi</tp:taxon-name-part></tp:taxon-name></italic> lack the information of the precise data of the collecting sites and dates, which however were indicated in the original descriptions (<xref ref-type="bibr" rid="B11">Nakahara 1955a</xref>, <xref ref-type="bibr" rid="B12">b</xref>).</p>
          <!--PageBreak-->
          <p>It is interesting that this species is polymorphic. Some individuals (See Figs <xref ref-type="fig" rid="F8">28</xref>, <xref ref-type="fig" rid="F8">30</xref>) have dark wings, while some individuals (See Figs <xref ref-type="fig" rid="F8">29</xref>, <xref ref-type="fig" rid="F8">31</xref>) have much paler wings. Therefore, it is difficult to identify the Taiwanese <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> species only based on external morphology because of its significant intraspecific variations. Possibly, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">formosanus</tp:taxon-name-part></tp:taxon-name></italic> and <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">pallidus</tp:taxon-name-part></tp:taxon-name></italic> are individual variations of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">taiwanensis</tp:taxon-name-part></tp:taxon-name></italic>, which, however, needs further clarification.</p>
          <p>Considering the distribution, this species is firstly recorded in Iriomote Island, which is the southernmost island of Ryukyus, Japan.</p>
          <fig id="F4" position="float" orientation="portrait">
            <object-id content-type="doi">10.3897/dez.61.8793.figures17-20</object-id>
            <object-id content-type="zenodo_dep_id">959865</object-id>
            <object-id content-type="arpha">BB6A322A-41A5-5284-AFF1-AEFA1ECBD3F8</object-id>
            <label>Figures 17–20.</label>
            <caption>
              <p><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">taiwanensis</tp:taxon-name-part></tp:taxon-name></italic> Banks. <bold>17.</bold> Male genitalia, dorsal view; <bold>18.</bold> Male genitalia, ventral view; <bold>19.</bold> Male genitalia, lateral view; <bold>20.</bold> Male ectoproct, caudal view. Scale bars: 0.5 mm.</p>
            </caption>
            <graphic xlink:href="dez-61-141-g004.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_34980.jpg">
              <uri content-type="original_file">https://binary.pensoft.net/fig/34980</uri>
            </graphic>
          </fig>
        </tp:treatment-sec>
      </tp:taxon-treatment>
      <tp:taxon-treatment>
        <tp:treatment-meta>
          <kwd-group>
            <label>Taxon classification</label>
            <kwd>
              <named-content content-type="kingdom" xlink:type="simple">Animalia</named-content>
            </kwd>
          </kwd-group>
        </tp:treatment-meta>
        <tp:nomenclature>
          <tp:taxon-name><object-id content-type="arpha">14BAE131-7479-5D60-8419-3F8DDFBA68CB</object-id>
            <tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part>
            <tp:taxon-name-part taxon-name-part-type="species">formosanus</tp:taxon-name-part>
          </tp:taxon-name>
          <tp:taxon-authority>(Okamoto &amp; Kuwayama, 1920)</tp:taxon-authority>
          <xref ref-type="fig" rid="F1">Figs 4</xref>
          <xref ref-type="fig" rid="F5">, 21</xref>
          <xref ref-type="fig" rid="F5">–22</xref>
          <tp:nomenclature-citation-list>
            <tp:nomenclature-citation>
              <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Lidar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">formosanus</tp:taxon-name-part></tp:taxon-name>
              <comment> Okamoto &amp; Kuwayama, 1920: 341. Type locality: China (Taiwan: Jiayi).</comment>
            </tp:nomenclature-citation>
          </tp:nomenclature-citation-list>
        </tp:nomenclature>
        <tp:treatment-sec sec-type="Diagnosis">
          <title>Diagnosis.</title>
          <p>This species is characterized by the forewings with numerous brown markings, and the basal part of bursa copulatrix, which is subtrapezoidal, with anteromedial portion slightly prominent and sclerotized in ventral view.</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Description">
          <title>Description.</title>
          <p>Female. Body length 8.5 mm; forewing length 14.0 mm, hindwing length 12.0 mm.</p>
          <p>Head yellowish brown, with pale yellow setose tubercles. Compound eyes blackish brown. Antenna with ca. 25 segments, pale yellowish brown, medial flagellomeres much longer than those on both ends, distal eight flagellomeres short, nearly moniliform.</p>
          <p>Prothorax pale yellowish brown, pronotum dark brown, with eight yellowish tubercles clothed with yellowish hairs; mesothorax and metathorax pale yellowish brown. Legs pale yellowish brown, with each segment blackish brown at tip. Wings hyaline, slightly pale brown, with numerous brownish spots. Forewing ~2.5 times as long as wide, densely spotted, most spots expanded and fused with each other, with markings on costal area and proximal half much darker, arranging as transversely arcuate pattern, an immaculate area present distal to median nygmata; two nygmata present on proximal and median portion of forewing, median one much larger than proximal one. Hindwing slightly paler than forewing, with similar marking pattern; one nygma present at middle. Veins brown. Forewing with 13 crossveins between Sc and R; Sc just touching R in pterostigmatic region; 12 crossveins between R and Rs; Rs with five main branches; MP with two main branches. Hindwing subcostal area slightly widened on distal half, with four crossveins; Rs with six main branches.</p>
          <p>Abdomen yellow, pregenital segments dorsally brown. Ovipositor yellowish brown, much longer than abdomen. Seventh sternite in lateral view subtrapezoidal, with slightly convex posterior margin in ventral view. Eighth abdominal segment without subgenitale. Bursa copulatrix with colleterial gland tubular and elongate, strongly curved medially; basal part of bursa copulatrix sac-like in lateral view, subtrapezoidal in ventral view with anteromedial portion slightly prominent and sclerotized; bursal accessory gland not observed. Ectoproct rather small, ovoid.</p>
          <p>Male. Unknown.</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Material examined">
          <title>Material examined.</title>
          <p>Holotype f#, “Mt. Ari [<named-content content-type="dwc:verbatimCoordinates">23°26′N, 120°46′E</named-content>], Taiwan, 2.IV.1907. S. Matsumur[a] leg./<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Lidar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">formosanus</tp:taxon-name-part></tp:taxon-name></italic> n. sp. 1920 Det. HANJIRO OKAMOTO SATORO KUWAYAMA (HOLOTYPE)/<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Lidar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">formosanus</tp:taxon-name-part></tp:taxon-name></italic> OKAMOTO et KUWAYAMA 1920 Holotype ♀” (SEHU).</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Distribution">
          <title>Distribution.</title>
          <p>China (Taiwan).</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Remarks">
          <title>Remarks.</title>
          <p>This species was originally described based on a single female by <xref ref-type="bibr" rid="B14">Okamoto and Kuwayama (1920)</xref>. Due to the phenomenon that remarkable intraspecific variation of wing marking patterns is present in <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="family">Dilaridae</tp:taxon-name-part></tp:taxon-name> (see <xref ref-type="bibr" rid="B22">Zhang et al. 2014b</xref> and the remarks on <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">taiwanensis</tp:taxon-name-part></tp:taxon-name></italic>), it is difficult to clarify the specific identity of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">formosanus</tp:taxon-name-part></tp:taxon-name></italic> based on the external morphology with the other Taiwanese <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> species, of which the females are unknown. Nevertheless, it is highly possible that <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">formosanus</tp:taxon-name-part></tp:taxon-name></italic> is conspecific with one of the other three <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> species from Taiwan, i.e. <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">pallidus</tp:taxon-name-part></tp:taxon-name></italic>, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">taiwanensis</tp:taxon-name-part></tp:taxon-name></italic>, and <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">insularis</tp:taxon-name-part></tp:taxon-name></italic>. This may be clarified by DNA barcoding, when fresh material is available.</p>
          <fig id="F5" position="float" orientation="portrait">
            <object-id content-type="doi">10.3897/dez.61.8793.figures21-22</object-id>
            <object-id content-type="zenodo_dep_id">959867</object-id>
            <object-id content-type="arpha">CFF8CE0D-97A9-5742-AEFE-FD52E9B72096</object-id>
            <label>Figures 21–22.</label>
            <caption>
              <p><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">formosanus</tp:taxon-name-part></tp:taxon-name></italic> (Okamoto &amp; Kuwayama). <bold>21.</bold> Female genitalia, lateral view; <bold>22.</bold> Female genitalia, ventral view. Scale bars: 0.5 mm.</p>
            </caption>
            <graphic xlink:href="dez-61-141-g005.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_34981.jpg">
              <uri content-type="original_file">https://binary.pensoft.net/fig/34981</uri>
            </graphic>
          </fig>
        </tp:treatment-sec>
      </tp:taxon-treatment>
      <tp:taxon-treatment>
        <tp:treatment-meta>
          <kwd-group>
            <label>Taxon classification</label>
            <kwd>
              <named-content content-type="kingdom" xlink:type="simple">Animalia</named-content>
            </kwd>
          </kwd-group>
        </tp:treatment-meta>
        <tp:nomenclature>
          <tp:taxon-name><object-id content-type="arpha">115C0E54-DA51-5141-AA67-86D2532B539C</object-id>
            <tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part>
            <tp:taxon-name-part taxon-name-part-type="species">pallidus</tp:taxon-name-part>
          </tp:taxon-name>
          <tp:taxon-authority>Nakahara, 1955</tp:taxon-authority>
          <xref ref-type="fig" rid="F1">Figs 5</xref>
          <xref ref-type="fig" rid="F6">, 23</xref>
          <tp:nomenclature-citation-list>
            <tp:nomenclature-citation>
              <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">pallidus</tp:taxon-name-part></tp:taxon-name>
              <comment> Nakahara, 1955b: 140. Type locality: China (Taiwan: Tattaka).</comment>
            </tp:nomenclature-citation>
          </tp:nomenclature-citation-list>
        </tp:nomenclature>
        <tp:treatment-sec sec-type="Diagnosis">
          <title>Diagnosis.</title>
          <p>This species is characterized by the forewings with no obvious markings, and the male ectoproct in dorsal view with a subrectangular projection terminating in three sharply pointed processes of about equal length.</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Description">
          <title>Description.</title>
          <p>Male. Body length 3.5 mm; forewing length 10.5 mm, hindwing length 8.5 mm.</p>
          <p>Head yellowish brown, with pale yellow setose tubercles. Compound eyes blackish brown. Antenna with ca. 24 segments, pale yellowish brown, pedicel with brown annular stripes, flagellum unipectinate on most flagellomeres, medial branches much longer than those on both ends, longest branch nearly 3.0 times as long as relevant flagellomere, but branch of 1st flagellomere short and dentate, distal eight flagellomeres simple.</p>
          <p>Prothorax pale yellowish brown, pronotum yellowish brown, with anterior margin and posterolateral corners pale yellow, medially with a pair of ovoid markings; mesothorax pale yellowish brown, mesonotum dark brown on anterior and lateral margins; metanotum pale yellowish brown, slight darker on lateral margins. Legs pale yellowish brown, femora blackish brown at tip. Wings hyaline, slightly yellowish brown. Forewing ~2.3 times as long as wide, with numerous very indistinct markings, and with a yellowish brown spot on median nygma; two nygmata present on proximal and median portion of forewing, median one much larger than proximal one. Hindwing ~2.1 times as long as wide, slightly paler than forewing; one nygma present at middle. Veins pale brown. Forewing with trichosors present along wing margin between R and CuP; Rs with four main branches; MP with two main branches; two gradate crossveins present at middle. Hindwing with trichosors present along wing margin between R and CuA; Rs with three main branches.</p>
          <p>Abdomen pale yellowish brown, pregenital segments dorsally brown. Ectoproct in dorsal view with a subrectangular projection which terminating in three sharply pointed processes of about equal length, posteroventrally with a pair of large, bifid, unguiform projections and a subrectangular, feebly sclerotized projection. Ninth gonocoxite with anterior half slightly inflated and with posterior half slenderly elongate and strongly incurved; <!--PageBreak-->tenth gonocoxite slenderly elongate, with incurved base and spinous tip, submedially with a lobe connecting to ninth gonocoxite; gonarcus beam-shaped, laterally connecting to base of ninth gonocoxites. Hypandrium internum unknown.</p>
          <p>Female. Unknown.</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Material examined">
          <title>Material examined.</title>
          <p>Holotype ♂, “Formosa, T. Kano/<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">pallidus</tp:taxon-name-part></tp:taxon-name></italic> n.sp. (Type) W. NAKAHARA/<italic>pallidus</italic> n. sp./Waro Nakahara Collection II/NSMT-I-Nr No. 4301/Type of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">pallidus</tp:taxon-name-part></tp:taxon-name></italic>?” (NSMT).</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Distribution">
          <title>Distribution.</title>
          <p>China (Taiwan).</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Remarks">
          <title>Remarks.</title>
          <p>This species is known only from the holotype male, whose genitalia have been unfortunately lost. The present redescription of the male genitalia is based on the original illustration from <xref ref-type="bibr" rid="B12">Nakahara (1955b)</xref>. Considering the wing marking patterns, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">pallidus</tp:taxon-name-part></tp:taxon-name></italic> has very pale wings without distinct dark markings, while the other Taiwanese species of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> have much darker markings on forewings. Considering the male genitalia, it is obvious that <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">pallidus</tp:taxon-name-part></tp:taxon-name></italic> is closely related to <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">taiwanensis</tp:taxon-name-part></tp:taxon-name></italic> by having similar male gonocoxite complexes 9, 10, 11. However, in <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">pallidus</tp:taxon-name-part></tp:taxon-name></italic> the male ectoproct differs in some details, e.g. the presence of a rectangular dorsomedian projection, and the presence of a pair of posteroventral lobes each with two widely separated claw-like projections, from <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">taiwanensis</tp:taxon-name-part></tp:taxon-name></italic>. Nevertheless, the specific identity of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">pallidus</tp:taxon-name-part></tp:taxon-name></italic> needs further clarification when more materials will be available.</p>
          <fig id="F6" position="float" orientation="portrait">
            <object-id content-type="doi">10.3897/dez.61.8793.figures23</object-id>
            <object-id content-type="zenodo_dep_id">959869</object-id>
            <object-id content-type="arpha">1C143D88-D653-567E-8A9D-059543228CD5</object-id>
            <label>Figures 23.</label>
            <caption>
              <p><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">pallidus</tp:taxon-name-part></tp:taxon-name></italic> Nakahara, male genitalia, ventral view. Reproduced from <xref ref-type="bibr" rid="B12">Nakahara (1955b)</xref>.</p>
            </caption>
            <graphic xlink:href="dez-61-141-g006.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_34982.jpg">
              <uri content-type="original_file">https://binary.pensoft.net/fig/34982</uri>
            </graphic>
          </fig>
        </tp:treatment-sec>
      </tp:taxon-treatment>
      <tp:taxon-treatment>
        <tp:treatment-meta>
          <kwd-group>
            <label>Taxon classification</label>
            <kwd>
              <named-content content-type="kingdom" xlink:type="simple">Animalia</named-content>
            </kwd>
          </kwd-group>
        </tp:treatment-meta>
        <tp:nomenclature>
          <tp:taxon-name><object-id content-type="arpha">DDA0CA46-7F7F-5648-A6E6-02BB05B0DFBE</object-id>
            <tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part>
            <tp:taxon-name-part taxon-name-part-type="species">insularis</tp:taxon-name-part>
            <object-id content-type="zoobank" xlink:type="simple">http://zoobank.org/721BA7E1-F9DF-4967-822B-E46BE6AA0C96</object-id>
          </tp:taxon-name>
          <tp:taxon-authority>Zhang, Liu &amp; U. Aspöck</tp:taxon-authority>
          <tp:taxon-status>sp. n.</tp:taxon-status>
          <xref ref-type="fig" rid="F1">Figs 6</xref>
          <xref ref-type="fig" rid="F7">, 24</xref>
          <xref ref-type="fig" rid="F7">–27</xref>
          <!--PageBreak-->
        </tp:nomenclature>
        <tp:treatment-sec sec-type="Diagnosis">
          <title>Diagnosis.</title>
          <p>This species is characterized by the forewings with numerous yellowish brown stripes, and the male gonarcus/ninth gonocoxite complex with rather inflated ninth gonocoxites and slenderly elongate tenth gonocoxites which are angulately curved anteriorly.</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Description">
          <title>Description.</title>
          <p>Male. Body length 3.7–5.5 mm; forewing length 6.0–8.9 mm, hindwing length 5.2–7.5 mm.</p>
          <p>Head yellowish brown, with pale yellow setose tubercles. Compound eyes blackish brown. Antenna with ca. 28 segments, pale yellowish brown, pedicel with brown annular stripes, flagellum unipectinate on most flagellomeres, medial branches much longer than those on both ends, longest branch nearly 7.0 times as long as relevant flagellomere, but branch of 1st flagellomere short and dentate, distal eight flagellomeres simple.</p>
          <p>Prothorax pale yellow, pronotum pale yellowish brown, with anterior margin and posterolateral corners pale yellow, medially with a pair of ovoid markings; mesothorax pale yellowish brown, mesonotum yellowish brown on anterior and lateral margins; metanotum yellowish brown, slightly darker on lateral margins. Legs pale yellowish brown, femora blackish brown at tip. Wings hyaline, slightly yellowish brown, with numerous yellowish brown stripes. Forewing ~2.2 times as long as wide, proximal stripes slightly darker, arranging as transversely arcuate pattern, an immaculate area present distal to median nygma; two nygmata present on proximal and median portion, median one much larger than proximal one. Hindwing ~2.1 times as long as wide, slightly paler than forewing, with similar marking pattern; one nygma present at middle. Veins pale brown. Forewing with trichosors present along wing margin between R and CuP; Sc just touching R in pterostigmatic region, terminally leaving several weak veinlets; Rs with four or five main branches; MP with two main branches; two gradate crossveins present at middle. Hindwing with trichosors present along wing margin between R and CuA; Rs with four main branches.</p>
          <p>Abdomen pale yellowish brown, pregenital segments dorsally dark brown. Ninth tergite in dorsal view with an arcuate anterior incision and a nearly V-shaped posterior incision, leaving a narrow median portion and a pair of subtriangular hemitergites, which are obtuse distally and densely haired; posteromedial portion protruding into a projection; in lateral view broad, with straight ventral margin and arcuate posterior margin. Ninth sternite obviously shorter than ninth tergite, arcuately convex posteriad. Ectoproct in dorsal view nearly truncate distally, with two pairs of claw-like projections, in ventral view posteroventrally with two pairs of unguiform projections and a pair of short, feebly sclerotized, digitiform projections. Ninth gonocoxite rather short and robust with acutely tapering tip; tenth gonocoxite slenderly elongate, slightly longer than ninth gonocoxite, angulately curved anteriorly, posteriorly with obtuse tip, medially with a lobe connecting to ninth gonocoxite; gonarcus slenderly beam-shaped, laterally connecting to base of ninth gonocoxites. Hypandrium internum subtriangular, with lateral margins slightly arcuate.</p>
          <p>Female. Unknown.</p>
          <!--PageBreak-->
          <!--PageBreak-->
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Type materials">
          <title>Type materials.</title>
          <p>Holotype ♂, “CHINA, Hainan, Mt. Wuzhishan [<named-content content-type="dwc:verbatimCoordinates">18°46’N, 109°30’E</named-content>], 1800 m, 23.V.1964, Sikong Liu” (CAU). Paratypes 1♂, CHINA, Hainan, Baisha, Bawangling [<named-content content-type="dwc:verbatimCoordinates">19°08’N, 109°15’E</named-content>], 1300 m, 24.V.2007 (CAU); 1♂, CHINA, Hainan, Lingshui, Mt. Diaoluoshan [<named-content content-type="dwc:verbatimCoordinates">18°39’N, 109°54’E</named-content>], 27.III.1964, Sikong Liu (CAU); 1♂, CHINA, Taiwan, Nantou [<named-content content-type="dwc:verbatimCoordinates">23°55’N, 120°40’E</named-content>], 10.VII.1961, T. Shirogu (NSMT).</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Distribution">
          <title>Distribution.</title>
          <p>China (Hainan, Taiwan).</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Etymology">
          <title>Etymology.</title>
          <p>The specific epithet “<italic>insularis</italic>” refers to the fact that the new species is endemic to the islands of East Asia: Hainan and Taiwan. It is an adjective in the nominative masculine singular.</p>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Remarks">
          <title>Remarks.</title>
          <p>This species can be easily distinguished from the other <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> species from Taiwan by the male ninth tergite with a posteromedial projection, the male gonocoxite complex 9, 10, 11 with rather inflated and short ninth gonocoxites, and the male ectoproct distally with two pairs of claw-like projections. First, we discovered this new species in several localities on Hainan Island, but subsequently found one specimen of this species in Taiwan, which indicates that this species might be diverged much earlier than the other Taiwanese <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> species when these two large islands were parts of the East Asian mainland.</p>
          <fig id="F7" position="float" orientation="portrait">
            <object-id content-type="doi">10.3897/dez.61.8793.figures24-27</object-id>
            <object-id content-type="zenodo_dep_id">959871</object-id>
            <object-id content-type="arpha">AD7C3CBD-1B9D-59AA-BED6-EBB2D42FDB8C</object-id>
            <label>Figures 24–27.</label>
            <caption>
              <p><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">insularis</tp:taxon-name-part></tp:taxon-name></italic> sp. n. <bold>24.</bold> Male genitalia, dorsal view; <bold>25.</bold> Male genitalia, ventral view; <bold>26.</bold> Male genitalia, lateral view; <bold>27.</bold> Male ectoproct, caudal view. Scale bars: 0.5 mm.</p>
            </caption>
            <graphic xlink:href="dez-61-141-g007.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_34983.jpg">
              <uri content-type="original_file">https://binary.pensoft.net/fig/34983</uri>
            </graphic>
          </fig>
          <fig id="F8" position="float" orientation="portrait">
            <object-id content-type="doi">10.3897/dez.61.8793.figures28-31</object-id>
            <object-id content-type="zenodo_dep_id">959873</object-id>
            <object-id content-type="arpha">9B8A5A2A-97C3-50F0-BFC6-FD278D594BFA</object-id>
            <label>Figures 28–31.</label>
            <caption>
              <p>Individual variations of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> spp. <bold>28, 30.</bold><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">taiwanensis</tp:taxon-name-part></tp:taxon-name></italic> Banks, male with dark wings; <bold>29, 31.</bold> same species, male with bright wings. Scale bars: 1.0 mm.</p>
            </caption>
            <graphic xlink:href="dez-61-141-g008.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_34984.jpg">
              <uri content-type="original_file">https://binary.pensoft.net/fig/34984</uri>
            </graphic>
          </fig>
          <fig id="F9" position="float" orientation="portrait">
            <object-id content-type="doi">10.3897/dez.61.8793.figure32</object-id>
            <object-id content-type="zenodo_dep_id">959875</object-id>
            <object-id content-type="arpha">7B6E7579-CBBD-50C5-B83B-4349266C65A4</object-id>
            <label>Figure 32.</label>
            <caption>
              <p>Geographic distribution of the species of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic>. ▲ <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">formosanus</tp:taxon-name-part></tp:taxon-name></italic> (Okamoto &amp; Kuwayama);  <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">hikosanus</tp:taxon-name-part></tp:taxon-name></italic> Nakahara; ● <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">insularis</tp:taxon-name-part></tp:taxon-name></italic> sp. n.; ■ <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">japonicus</tp:taxon-name-part></tp:taxon-name></italic> McLachlan; ▼ <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">pallidus</tp:taxon-name-part></tp:taxon-name></italic> Nakahara;  <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">taiwanensis</tp:taxon-name-part></tp:taxon-name></italic> Banks.</p>
            </caption>
            <graphic xlink:href="dez-61-141-g009.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_34985.jpg">
              <uri content-type="original_file">https://binary.pensoft.net/fig/34985</uri>
            </graphic>
          </fig>
          <!--PageBreak-->
        </tp:treatment-sec>
      </tp:taxon-treatment>
    </sec>
    <sec sec-type="Discussion" id="sec5">
      <title>Discussion</title>
      <p>Among the presently described <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> species, the two Taiwanese species (i.e. <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">pallidus</tp:taxon-name-part></tp:taxon-name></italic> and <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">taiwanensis</tp:taxon-name-part></tp:taxon-name></italic>) and the two Japanese species (i.e. <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">hikosanus</tp:taxon-name-part></tp:taxon-name></italic> and <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">japonicus</tp:taxon-name-part></tp:taxon-name></italic>) are morphologically similar and probably closely related because of the similar structure of the male gonocoxite complex 9, 10, 11, which has a strongly ventrally curved gonocoxite 9 and a proximally bifurcated gonocoxite 10. Based on these characters, the above four insular species appear to be closely related to the mainland species <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">septentrionalis</tp:taxon-name-part></tp:taxon-name></italic> Navás, 1912, which is distributed in northeastern Asia (northeastern China, Korea, and Russia Far East) (<xref ref-type="bibr" rid="B7">Monserrat 1988a</xref>, <xref ref-type="bibr" rid="B21">Zhang et al. 2014a</xref>). Whether the common ancestor of these four species from the islands invaded the islands and diversified there afterwards or whether the island species result from several invaders remains open at present. Hypotheses on the separation of the islands offshore Eastern Asia have been discussed in the context of several taxa, e.g. <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order">Raphidioptera</tp:taxon-name-part></tp:taxon-name> (<xref ref-type="bibr" rid="B5">Liu et al. 2010</xref>). The subsequent southern dispersal of the ancestral species from Japan through Ryukyus to Taiwan might account for further separation and the modern distribution of these four insular species.</p>
      <p><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">insularis</tp:taxon-name-part></tp:taxon-name></italic> sp. n. is remarkably different from the other insular species from Japan and Taiwan, and it seems to be somewhat related to the <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Dilar</tp:taxon-name-part></tp:taxon-name></italic> species ranging in the Himalayas (<xref ref-type="bibr" rid="B22">Zhang et al. 2014b</xref>) by having the posteromedial projection of male ninth tergite, suggesting that the speciation of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Dilar">D.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species">insularis</tp:taxon-name-part></tp:taxon-name></italic> sp. n. might be correlated with a different vicariance event, possibly the separation of Hainan and Taiwan from the southeastern Chinese mainland.</p>
    </sec>
  </body>
  <back>
    <ack>
      <title>Acknowledgements</title>
      <p>We thank Dr. Utsugi Jinbo (NSMT, Tsukuba), Dr. Yukinobu Nakatani (NIAES, Tsukuba), Dr. Kazunori Yoshizawa (SEHU, Sapporo), Dr. John D. Oswald (College Station, Texas), Dr. Oliver S. Flint, Jr (NMNH, Washington), Dr. Benjamin W. Price (BMNH, London), and Dipl. Biol. Hella Wendt (MNB, Berlin) for processing the loan of specimens for our study. This research was supported by the National Natural Science Foundation of China (Nos. 31322051 and 31320103902).</p>
    </ack>
    <ref-list>
      <title>References</title>
      <ref id="B1">
        <mixed-citation xlink:type="simple"><person-group><name name-style="western"><surname>Aspöck</surname><given-names>H</given-names></name><name name-style="western"><surname>Hölzel</surname><given-names>H</given-names></name><name name-style="western"><surname>Aspöck</surname><given-names>U</given-names></name></person-group> (<year>2001</year>) <article-title>Kommentierter Katalog der Neuropterida (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="class">Insecta</tp:taxon-name-part></tp:taxon-name>: <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order">Raphidioptera</tp:taxon-name-part></tp:taxon-name>, <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order">Megaloptera</tp:taxon-name-part></tp:taxon-name>, <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order">Neuroptera</tp:taxon-name-part></tp:taxon-name>) der Westpaläarktis.</article-title><source>Denisia</source><volume>2</volume>: <fpage>1</fpage>–<lpage>606</lpage>.</mixed-citation>
      </ref>
      <ref id="B2">
        <mixed-citation xlink:type="simple"><person-group><name name-style="western"><surname>Aspöck</surname><given-names>U</given-names></name><name name-style="western"><surname>Aspöck</surname><given-names>H</given-names></name></person-group> (<year>2008</year>) <article-title>Phylogenetic relevance of the genital sclerites of Neuropterida (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="class">Insecta</tp:taxon-name-part></tp:taxon-name>: Holometabola).</article-title><source>Systematic Entomology</source><volume>33</volume>: <fpage>97</fpage>–<lpage>127</lpage>. doi: <ext-link ext-link-type="doi" xlink:href="10.1111/j.1365-3113.2007.00396.x">10.1111/j.1365-3113.2007.00396.x</ext-link></mixed-citation>
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    <fn-group>
      <fn id="FN1">
        <p>Dilar formosanus (Okamoto &amp; Kuwayama) is not included in this key because the male of this species is unknown. Moreover, the specific identity of D. formosanus is still unclear (see remarks of D. formosanus).</p>
      </fn>
    </fn-group>
  </back>
</article>
