This species is easily confused with several species having unicolourous reddish brown body in this subgenus, but can be recognized by the stout penis guide with a hook-like appendage at apex in lateral view (Fig. 1h).

TL: 2.31–2.43 mm, TW: 1.53–1.67 mm, TH: 0.98–1.04 mm, TL/TW: 1.46–1.51, PL/PW: 0.56–0.57, EL/EW: 1.13–1.16, HW/PW: 0.59–0.62, PW/EW: 0.73–0.74.

Body elongate oval, moderately convex, entirely reddish brown, dorsum covered with white pubescence (Fig. 1a–c).

Head with fine frontal punctures, as large as eye facets, 1.0–2.0 diameters apart. Eyes densely faceted, interocular distance 0.51 times head width. Pronotal punctures as large as those on frons, 2.0–3.0 diameters apart. Surface of elytra with punctures larger than those on pronotum, separated by 2.5–3.0 diameters. Prosternal process trapezoidal, 2 times as long as its width at base; with lateral carinae distinct, extending to anterior margin of prosternum, distinctly convergent anteriorly. Abdominal postcoxal lines weakly recurved and distinctly incomplete laterally (Fig. 1d), reaching 6/7 length of abdominal ventrite 1, area enclosed by lines coarsely punctate, broadly smooth along line. Abdominal ventrite 5 in male with apex truncate.

Male genitalia: penis stout, unevenly sinuated (Fig. 1e). Penis capsule highly sclerotized with small inner arm and large outer arm (Fig. 1e). Apex of penis strongly curved with distinctly membranous appendage (Fig. 1f). Tegmen extremely stout (Fig. 1g–h) with penis guide narrowest at base, gradually expanding to a spade-like apex bearing with a digitations in ventral view (Fig. 1g); in lateral view, penis guide strongly raised at middle with a hook-like appendage at apex (Fig. 1h). Parameres slender with rounded apex, slightly longer than penis guide, densely covered with two groups of long setae at apices (Fig. 1h).

Female unknown.

Holotype: male, No. SCAU (E) 11346, China: Gansu: Dongshan, Xinglongshan National Nature Reserve, Yuzhong County, Lanzhou City, 35°47.83’N, 104°3.93’E, ca 2300 m, 3. VIII. 2007, Liang JB leg. Paratypes (3): 3♂ with same data as holotype.

China (Gansu).

The species name is derived from the Latin adjective hastatus referring to its penis guide with halberd-like apex in ventral view.

This species is most similar to Scymnus (Parapullus) alishanensis Pang & Yu, 1993 in general appearance and male genitalia, but can be distinguished from it by having parameres as long as half length of penis guide (Fig. 2h) and penis guide constricted at middle part in ventral view (Fig. 2g). In S. (P.) alishanensis, its parameres are longer than half length of penis guide and penis guide constricted near middle part in ventral view.

TL: 2.20–2.48 mm, TW: 1.60–1.71 mm, TH: 1.06–1.18 mm, TL/TW: 1.38–1.45, PL/PW: 0.54–0.55, EL/EW: 1.06–1.09, HW/PW: 0.59–0.60, PW/EW: 0.70–0.75.

Body elongate oval, moderately convex, entirely reddish brown, dorsum covered with white pubescence (Fig. 2a–c).

Head with fine frontal punctures, slightly smaller than eye facets, 0.5–1.0 diameter apart. Eyes densely faceted, interocular distance 0.49 times head width. Pronotal punctures larger than those on frons, 1.0–2.0 diameters apart. Surface of elytra with punctures much larger than those on pronotum, separated by 1.0–2.0 diameters. Prosternal process trapezoidal, 2 times as long as its width at base; with lateral carinae distinct, extending to anterior margin of prosternum, slightly convergent anteriorly. Abdominal postcoxal lines weakly recurved and distinctly incomplete laterally (Fig. 2d), reaching 6/7 length of abdominal ventrite 1, area enclosed by lines finely punctate, narrowly smooth along line. Abdominal ventrite 5 in both sexes with apex truncate.

Male genitalia: penis stout (Fig. 2e). Penis capsule deeply emarginated medially with small inner arm and large outer arm. Apex of penis slender, forming a hook-shaped (Fig. 2f). Tegmen stout (Figs 2g–h) with penis guide slightly constricted at middle part, expanding toward apex, then tapering abruptly to blunt apex in ventral view (Fig. 2g). Parameres narrow, as long as half length of penis guide, densely covered with two groups of long setae at apices (Fig. 2h).

Female similar to male except for sexual characters.

Holotype: male, No. SCAU (E) 11532, China: Tianjin: Baxianshan National Nature Reserve, Ji County, 40°11.27’N, 117°33.19’E, ca 500 m, 14. VII. 2011, Chen XS leg. Paratypes (3): 1♂2♀ with same data as holotype.

China (Tianjin).

Etymology. The specific epithet refers to its type locality, Baxianshan National Nature Reserve, Tianjin.

This species is similar to Scymnus (Parapullus) parallelicus Chen, Ren & Wang, 2012 in general appearance and male genitalia, but can be distinguished from it by having strongly sinuated apex of penis with membranous appendage (Fig. 3f), the apical 1/4 length of parameres covered with several long setae at inner sides in lateral view (Fig. 3h), and the area enclosed by the abdominal postcoxal lines with coarse punctures (Fig. 3d).

TL: 2.21–2.38 mm, TW: 1.58–1.71 mm, TH: 1.01–1.16 mm, TL/TW: 1.39–1.40, PL/PW: 0.51–0.52, EL/EW: 1.08–1.12, HW/PW: 0.56, PW/EW: 0.72.

Body elongate oval, moderately convex, entirely reddish brown, dorsum covered with white pubescence (Fig. 3a–c).

Head with fine frontal punctures, as large as eye facets, 0.5–1.0 diameter apart. Eyes densely faceted, interocular distance 0.47 times head width. Pronotal punctures similar to those on frons, 1.0–2.0 diameters apart. Surface of elytra with punctures larger than those on pronotum, separated by 2.0–3.0 diameters. Prosternal process trapezoidal, 2.5 times as long as its width at base; with lateral carinae distinct, extending to anterior margin of prosternum, distinctly convergent anteriorly. Abdominal postcoxal lines weakly recurved and distinctly incomplete laterally (Fig. 3d), reaching 5/6 length of abdominal ventrite 1, area enclosed by lines coarsely punctate, narrowly smooth along line. Abdominal ventrite 5 in male with apex rounded.

Male genitalia: penis stout (Fig. 3e). Penis capsule with small inner arm and large outer arm. Apex of penis strongly sinuated with membranous appendage (Fig. 3f). Tegmen stout (Fig. 3g–h) with penis guide with parallel sides from base to 3/4 length, then tapering gradually to blunt apex in ventral view (Fig. 3g) and its apex slightly curved outwardly in lateral view (Fig. 3h). Parameres longer than half length of penis guide, densely covered with a group of long setae at apices, and its apical 1/4 length covered with several stout setae at inner sides (Fig. 3h).

Female externally similar to male but with abdominal ventrite 5 truncate apically.

Holotype: male, No. SCAU (E) 11534, China: Henan: Laojieling National Nature Reserve, Xixia County, Nanyang City, 33°37.63’N, 111°45.85’E, ca 1400 m, 9. VII. 2009, Chen XS leg. Paratypes (3): 1♂2♀ with same data as holotype.

China (Henan).

The specific epithet refers to its type locality, Laojieling National Nature Reserve, Henan Province.

This species is similar to Scymnus (Parapullus) parallelicus Chen, Ren & Wang, 2012 in general appearance and male genitalia, but can be separated from it by having slender, ring shaped apex of penis (Fig. 4f) and the abdominal postcoxal lines strongly recurved toward base of ventrite (Fig. 4d). It also resembles Scymnus (Parapullus) malleatus Chen, Ren & Wang, 2012, particularly the ring shaped apex of penis, but can be recognized from it by having narrow and short parameres (Fig. 4h) and the different shape of penis capsule.

TL: 2.28–2.34 mm, TW: 1.58–1.71 mm, TH: 1.06–1.15 mm, TL/TW: 1.37–1.44, PL/PW: 0.51–0.53, EL/EW: 1.07–1.13, HW/PW: 0.60–0.61, PW/EW: 0.67–0.71.

Body oval, moderately convex, entirely reddish brown, dorsum covered with white pubescence (Fig. 4a–c).

Head with fine frontal punctures, as large as eye facets, 1.0–2.0 diameters apart. Eyes densely faceted, interocular distance 0.49 times head width. Pronotal punctures larger than those on frons, 1.0–2.0 diameters apart. Surface of elytra with punctures larger than those on pronotum, separated by 2.0–3.0 diameters. Prosternal process trapezoidal, 2.5 times as long as its width at base; with lateral carinae distinct, extending to anterior margin of prosternum, slightly convergent anteriorly. Abdominal postcoxal lines strongly recurved and distinctly incomplete laterally (Fig. 4d), reaching 5/6 length of abdominal ventrite 1, area enclosed by lines finely punctate, broadly smooth along line. Abdominal ventrite 5 in both sexes with apex truncate.

Male genitalia: penis moderately stout (Fig. 4e). Penis capsule with small inner arm and large outer arm. Apex of penis slender, ring shaped (Fig. 4f). Tegmen stout (Fig. 4g–h) with penis guide parallel sided from base to 3/4 length, then tapering gradually to blunt apex in ventral view (Fig. 4g). Parameres stout with obliquely truncate apex, longer than half length of penis guide, densely covered with two groups of long setae at apices (Fig. 4h).

Female similar to male except for sexual characters.

Holotype: male, No. SCAU (E) 11527, China: Anhui: Hongshan, Yaoluoping National Nature Reserve, Yuexi County, 31°0.85’N, 116°8.67’E, ca 800 m, 30. IX. 2010, Wang XM leg. Paratypes (9): Anhui: 1♂ with same data as holotype. 1♂, Yaoluoping National Nature Reserve, Yuexi County, 30°58.95’N, 116°4.83’E, ca 1000 m, 28. IX. 2010, Li WJ leg. Tibet: 1♀, Cibagou National Nature Reserve, Chayu County, 28°33.69’N, 97°5.10’E, ca 2000 m, 13. X. 2007, Chen XS leg. 2♀, Xiachayu Town, Chayu County, 28°30.04’N, 97°0.84’E, ca 1800 m, 14. X. 2007, Chen XS leg. 1♂, Xiayadong Village, Yadong County, 28°29.29’N, 97°1.36’E, ca 2800 m, 29-30. IX. 2009, Chen XS leg. 1♀, Motuo County, Linzhi City, 29°19.30’N, 95°18.33’E, ca 760 m, 17. X. 2009, Chen XS leg. 1♀, Lebu Village, Cuona County, 27°48.63’N, 91°44.98’E, ca 2400 m, 24. X. 2011, Li WJ leg. 1♂, Mama Village, Cuona County, 27°47.44’N, 91°46.56’E, ca 2800 m, 25. X. 2011, Huo LZ leg.

China (Anhui, Tibet).

The specific epithet refers to its penis with ring shaped apex.

This species is similar to Scymnus (Parapullus) solus Chen, Ren & Wang, 2012 in general appearance and male genitalia, but can be separated from it by having a papilla bearing a stout seta at the apical 1/3 length of paramere (Fig. 5h).

TL: 1.98–2.36 mm, TW: 1.39–1.66 mm, TH: 0.92–1.09 mm, TL/TW: 1.42, PL/PW: 0.50–0.55, EL/EW: 1.09–1.12, HW/PW: 0.59–0.60, PW/EW: 0.70–0.73.

Body elongate oval, moderately convex, dorsum covered with white pubescence (Fig. 5a–c). Head, antennae and mouthparts brown. Pronotum dark brown. Scutellum black. Elytra black with apical 1/4 brown. Prothoracic hypomeron and prosternum brown. Mesoventrite, metaventrite and elytral epipleura black. Legs brown.

Head with fine frontal punctures, as large as eye facets, 0.5–1.0 diameter apart. Eyes densely faceted, interocular distance 0.50 times head width. Pronotal punctures larger than those on frons, 1.5–3.0 diameters apart. Surface of elytra with punctures as large as those on pronotum, separated by 1.0–2.0 diameters. Prosternal process trapezoidal, 2.3 times as long as its width at base; with lateral carinae distinct, extending to anterior margin of prosternum, distinctly convergent anteriorly. Abdominal postcoxal lines weakly recurved and distinctly incomplete laterally (Fig. 5d), nearly extending to posterior margins of abdominal ventrite 1, area enclosed by lines finely punctate, narrowly smooth along line. Abdominal ventrite 5 in male with apex truncate.

Male genitalia: penis stout and long (Fig. 5e). Penis capsule with outer arm large and inner arm strongly curved. Apex of penis unevenly curved, forming a large hook (Fig. 5f). Tegmen stout (Fig. 5g–h) with penis guide narrowest at base, expanding toward apex, then tapering abruptly to blunt apex in ventral view (Fig. 5g). Parameres obliquely truncated apically, longer than half length of penis guide, densely covered with a group of long setae at apices, and its apical 1/3 length with a papilla bearing a stout seta on inner side (Fig. 5h).

Female externally similar to male but with abdominal ventrite 5 rounded apically.

Holotype: male, No. 20050319043, China: Guangdong: Xiaohuangshan, Nanling National Nature Reserve, Shaoguan City, 24°54.76’N, 113°2.83’E, ca 800 m, 1. X. 2004, Ren SX leg. Paratypes (12): Guangdong: 1♂ with same data as holotype except “Chen JJ leg”. Hunan: 1♂, Mangshan National Nature Reserve, Yizhang County, 24°58.83’N, 112°53.14’E, ca 700 m, 2. X. 2004, Ren SX leg. Tianjin: 3♂6♀, Baxianshan National Nature Reserve, Ji County, 40°11.27’N, 117°33.19’E, ca 500 m, 14. VII. 2011, Chen XS leg. Shanxi: 1♂, Lishan National Nature Reserve, Yuanqu County, 35°28.39’N, 112°9.45’E, ca 1200 m, 3. VIII. 2011, Chen XS leg.

China (Shanxi, Tianjin, Hunan, Guangdong).

The specific epithet refers to its paramere with a papilla on the inner side of the apical 1/3 length.

This species is similar to Scymnus (Parapullus) aduncatus Chen, Ren & Wang, 2012 in general appearance, but can be distinguished from it by having robust basal part of penis guide in lateral view (Fig. 6h) and the blunt apex in lateral view (Fig. 6g).

TL: 2.16 mm, TW: 1.46 mm, TH: 0.97 mm, TL/TW: 1.48, PL/PW: 0.52, EL/EW: 1.14, HW/PW: 0.63, PW/EW: 0.71.

Body elongate oval, moderately convex, dorsum covered with white pubescence (Figs 6a–c). Head, antennae and mouthparts brown. Pronotum yellowish brown. Scutellum black. Elytra black with apical 1/6 yellowish brown. Prothoracic hypomeron and prosternum yellowish brown. Mesoventrite, metaventrite and elytral epipleura black. Legs brown.

Head with fine frontal punctures, as large as eye facets, 1.0–2.0 diameters apart. Eyes densely faceted, interocular distance 0.49 times head width. Pronotal punctures larger than those on frons, 1.0–2.0 diameters apart. Surface of elytra with punctures similar to those on pronotum, separated by 2.0–3.0 diameters. Prosternal process trapezoidal, 2 times as long as its width at base; with lateral carinae distinct, extending to anterior margin of prosternum, slightly convergent anteriorly. Abdominal postcoxal lines weakly recurved and distinctly incomplete laterally (Fig. 6d), reaching 4/5 length of abdominal ventrite 1, area enclosed by lines finely punctate, broadly smooth along line. Abdominal ventrite 5 in male with apex truncate.

Male genitalia: penis slender (Fig. 6e). Penis capsule stout with outer arm large and inner arm strongly curved upwardly. Apex of penis simple (Fig. 6f). Tegmen stout (Figs 6g–h) with penis guide narrowest at base, expanding toward apex, then tapering abruptly to blunt apex in ventral view (Fig. 6g) and strongly curved outwardly in lateral view (Fig. 6h). Parameres slightly constricted at base, distinctly shorter than penis guide, densely covered with two groups of long setae at apices (Fig. 6h).

Female unknown.

Holotype: male, No. SCAU (E) 11388, China: Yunnan: Xujiaba, Ailaoshan National Nature Reserve, Jingdong County, 24°32.77’N, 101°01.69’E, ca 2500 m, 17. VIII. 2013, Chen XS leg.

China (Yunnan).

The specific epithet refers to its apex of paramere densely covered with two groups of long setae.

This species is similar to Scymnus (Parapullus) secula Yang, 1978 in general appearance and male genitalia, but can be separated from it by the brown pronotum with a large black marking (Fig. 7a–b) and the apex of penis stout, hook-shaped (Fig. 7f).

TL: 2.18 mm, TW: 1.46 mm, TH: 0.98 mm, TL/TW: 1.49, PL/PW: 0.46, EL/EW: 1.20, HW/PW: 0.57, PW/EW: 0.77.

Body elongate oval, slightly convex, dorsum covered with white pubescence (Figs 7a–c). Head, antennae and mouthparts dark brown. Pronotum brown with a large black marking. Scutellum black. Elytra black with apical 1/6 yellowish brown. Prothoracic hypomeron and prosternum brown. Mesoventrite, metaventrite and elytral epipleura black. Legs brown.

Head with fine frontal punctures, slightly larger than eye facets, 0.5–1.0 diameter apart. Eyes densely faceted, interocular distance 0.50 times head width. Pronotal punctures as large as those on frons, 1.0–2.0 diameters apart. Surface of elytra with punctures larger than those on pronotum, separated by 2.0–3.0 diameters. Prosternal process trapezoidal, 2 times as long as its width at base; with lateral carinae distinct, extending to anterior margin of prosternum, slightly convergent anteriorly. Abdominal postcoxal lines strongly recurved and distinctly incomplete laterally (Fig. 7d), reaching 5/6 length of abdominal ventrite 1, area enclosed by lines irregularly punctate, narrowly smooth along line. Abdominal ventrite 5 in male with apical margin shallowly emarginated medially.

Male genitalia: penis stout (Fig. 7e). Penis capsule with small inner arm and large outer arm. Apex of penis stout with membranous appendage, hook-shaped (Fig. 7f). Tegmen stout (Figs 7g–h) with penis guide narrowest at base, expanding toward apex, then tapering abruptly to blunt apex in ventral view (Fig. 7g) and its apex slightly curved outwardly in lateral view (Fig. 7h). Parameres slightly curved at base, distinctly shorter than penis guide, densely covered with two groups of long setae at apices (Fig. 7h).

Female unknown.

Holotype: male, No. SCAU (E) 11455, China: Taiwan: Shenmu Railway Station, Alishan Mountains, Jiayi County, 23°31.12’N, 120°48.37’E, ca 2200 m, 23. X. 2012, Chen XS leg.

China (Taiwan).

The specific epithet refers to its type locality, Shenmu Railway Station in Alishan Mountains, Taiwan.

This species is similar to Scymnus (Parapullus) nanlingicus Chen, Ren & Wang, 2012 in general appearance and male genitalia, but can be separated from it by having broad area enclosed by abdominal postcoxal lines (Fig. 8d) and the parameres narrowest at base, expanding toward apex in lateral view (Fig. 8h).

TL: 2.20–2.42 mm, TW: 1.52–1.67 mm, TH: 0.99–1.05 mm, TL/TW: 1.45, PL/PW: 0.51, EL/EW: 1.13–1.14, HW/PW: 0.56–0.58, PW/EW: 0.73–0.74.

Body oval, moderately convex, dorsum covered with white pubescence (Fig. 8a–c). Head, antennae and mouthparts brown. Pronotum yellowish brown. Scutellum dark brown. Elytra black with apical 1/4 yellowish brown. Prothoracic hypomeron and prosternum yellowish brown. Mesoventrite, metaventrite and elytral epipleura black. Legs brown.

Head with fine frontal punctures, as large as eye facets, 0.5–1.0 diameters apart. Eyes densely faceted, interocular distance 0.47 times head width. Pronotal punctures similar to those on frons, 1.0–2.0 diameters apart. Surface of elytra with punctures larger than those on pronotum, separated by 2.0–3.0 diameters. Prosternal process rectangular, 2 times as long as its width at base; with lateral carinae distinct, parallel, extending to anterior margin of prosternum. Abdominal postcoxal lines strongly recurved and distinctly incomplete laterally (Fig. 8d), reaching 7/8 length of abdominal ventrite 1, area enclosed by lines irregularly punctate, broadly smooth along line. Abdominal ventrite 5 in male with apex truncate.

Male genitalia: penis stout (Fig. 8e). Penis capsule with outer arm stout and inner arm strongly curved, hook-shaped. Apex of penis simple (Fig. 8f). Tegmen stout (Fig. 8g–h) with penis guide parallel sided from base to 4/5 length, then tapering abruptly to blunt apex in ventral view (Fig. 8g). Parameres narrowest at base, expanding toward apex, distinctly shorter than penis guide, densely covered with two groups of long setae at apices (Fig. 8h).

Female unknown.

Holotype: male, No. SCAU (E) 11531, China: Sichuan: Yanzigou, Gonggashan National Nature Reserve, Luding County, 29°41.92’N, 102°0.22’E, ca 2500 m, 28. IX. 2007, Chen XS leg. Paratypes (1): 1♂ with same data as holotype.

China (Sichuan).

The specific epithet refers to its type locality, Yanzigou Scenic Spots, Gonggashan National Nature Reserve, Sichuan Province.