Latest Articles from Deutsche Entomologische Zeitschrift Latest 100 Articles from Deutsche Entomologische Zeitschrift https://dez.pensoft.net/ Fri, 29 Mar 2024 13:12:57 +0200 Pensoft FeedCreator https://dez.pensoft.net/i/logo.jpg Latest Articles from Deutsche Entomologische Zeitschrift https://dez.pensoft.net/ New genus and species of lice in the Oxylipeurus-complex (Phthiraptera, Ischnocera, Philopteridae), with an overview of the distribution of ischnoceran chewing lice on galliform hosts https://dez.pensoft.net/article/111874/ Deutsche Entomologische Zeitschrift 71(1): 85-109

DOI: 10.3897/dez.71.111874

Authors: Daniel R. Gustafsson, Chunpo Tian, Mengjiao Ren, Zhu Li, Xiuling Sun, Fasheng Zou

Abstract: Here, we describe a new genus of lice (Phthiraptera, Ischnocera) in the Oxylipeurus-complex, parasitising galliform hosts in the genera Tragopan Cuvier, 1829. This genus, Pelecolipeurus gen. nov., is separated from other members of the complex by the unique shape of the male subgenital plate and stylus, the male genitalia and other characters. The only previously-known species in the genus is Lipeurus longus Piaget, 1880, which is here tentatively re-described as Pelecolipeurus longus (Piaget, 1880), based on specimens from a non-type host, Tragopan temminckii (Gray, 1831). In addition, we describe a new species, Pelecolipeurus fujianensis sp. nov., based on specimens from Tragopan caboti (Gould, 1857). An overview of the distribution patterns of ischnoceran lice on galliforms is presented, which suggests that host phylogeny, host biogeography and host biotope, as well as elevation of host range, may all be important factors that have structured louse communities on landfowl. We transfer the genus Afrilipeurus from the Oxylipeurus-complex to the Lipeurus-complex and include an emended key to the Oxylipeurus-complex.

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Research Article Tue, 12 Mar 2024 09:36:08 +0200
Integrated taxonomy, biology and biogeography of the Afrotropical genus Xyloctonus (Coleoptera, Curculionidae, Scolytinae) https://dez.pensoft.net/article/116185/ Deutsche Entomologische Zeitschrift 71(1): 67-84

DOI: 10.3897/dez.71.116185

Authors: Bjarte H. Jordal

Abstract: The peculiar Afrotropical bark beetle genus Xyloctonus Eichhoff, 1872 is revised and its biology described. Several unusual morphological features reflect adaptations to predator avoidance as they are highly exposed during mating externally on tree trunks and branches. Observations invariably indicate that males and females abandon the nest under bark at an early stage of progeny, the males already before eggs hatch, potentially engaging in subsequent additional matings. Most species have a clear preference for host plants in the plant family Sapotaceae. Although the genus is broadly distributed in forested parts of Africa, Madagascar and Mauritius, most species are found in the eastern part of this range. A Bayesian biogeographical analysis revealed a possible origin of the genus in Madagascar in the early Eocene, with subsequent colonisation of the southern African region in late Eocene. This contrasts with the closely-related xyloctonine genus Ctonoxylon Hagedorn, 1910, which is of western Congolian ancestry and more recently reached Madagascar multiple times during late Miocene. Two new species are described: Xyloctonus magnus sp. nov. from Madagascar and X. genieri sp. nov. from Burkina Faso. Synonyms are proposed for X. subcostatus Eggers, 1939 (= X. striatus Eggers, 1939) and X. scolytoides Eichhoff, 1872 (= X. latus Eggers, 1922). Identification to species is provided in a key illustrated with photographs of most species.

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Research Article Fri, 1 Mar 2024 14:53:57 +0200
Two new species of the genus Agramma (Hemiptera, Heteroptera, Tingidae) from small islands of Japan, with an illustrated key to the Japanese species of the genus https://dez.pensoft.net/article/108270/ Deutsche Entomologische Zeitschrift 71(1): 49-65

DOI: 10.3897/dez.71.108270

Authors: Jun Souma

Abstract: The present study describes two new species of the monocotyledon-feeding lace bugs of the genus Agramma Stephens, 1829 (Hemiptera, Heteroptera, Tingidae, Tinginae, Tingini) from small islands of Japan. The first is A. (A.) izuense sp. nov., which was recorded as A. (A.) japonicum (Drake, 1948) from Hachijo Island, the Izu Islands, in a previous study, and is considered an independent species here based on morphological characteristics and molecular data. The second is A. (A.) keramense sp. nov., which has a remarkable spineless head and was discovered from Aka and Geruma islands, Kerama Group, the Ryukyu Islands. Consequently, the following four species of Agramma were recognized in Japan: A. (A.) abruptifrons Golub, 1990, A. (A.) izuense sp. nov., A. (A.) japonicum, and A. (A.) keramense sp. nov. Only dozens of submacropterous morphs were confirmed in these two species in the present study, suggesting that both new species are flightless. In addition, an illustrated key for the identification of the four species from Japan and the host plant relationships of the two new species are provided.

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Research Article Tue, 30 Jan 2024 12:18:57 +0200
Forest leaf litter beetles of Taiwan: first DNA barcodes and first insight into the fauna https://dez.pensoft.net/article/112278/ Deutsche Entomologische Zeitschrift 71(1): 17-47

DOI: 10.3897/dez.71.112278

Authors: Fang-Shuo Hu, Emmanuel Arriaga-Varela, Gabriel Biffi, Ladislav Bocák, Petr Bulirsch, Albert František Damaška, Johannes Frisch, Jiří Hájek, Peter Hlaváč, Bin-Hong Ho, Yu-Hsiang Ho, Yun Hsiao, Josef Jelínek, Jan Klimaszewski, Robin Kundrata, Ivan Löbl, György Makranczy, Keita Matsumoto, Guan-Jie Phang, Enrico Ruzzier, Michael Schülke, Zdeněk Švec, Dmitry Telnov, Wei-Zhe Tseng, Lan-Wei Yeh, My-Hanh Le, Martin Fikáček

Abstract: We report the publication of 953 DNA barcodes of forest leaf litter beetles from central Taiwan, in total representing 334 species of 36 beetle families. This is the first bulk of data from the Taiwanese Leaf Litter beetles project focused on uncovering the under-explored diversity of leaf litter beetles across Taiwan. Based on these data, we provide the first records of the following taxa for Taiwan: family Sphindidae (genus Aspidiphorus Ziegler, 1821); tribes Trichonychini, Ctenistini, and Bythinoplectini (all Staphylinidae: Pselaphinae); genera Gyrelon Hinton, 1942, Thyroderus Sharp, 1885, Cautomus Sharp, 1885 (all Cerylonidae), Dermatohomoeus Hlisnikovský, 1963 (Leiodidae), Paraploderus Herman, 1970 (Staphylinidae: Oxytelinae), Thinocharis Kraatz, 1859 (Staphylinidae: Paederinae), Cephennodes Reitter, 1884, Napoconnus Franz, 1957 (both Staphylinidae: Scydmaeninae), Bicava Belon, 1884 (Latridiidae), Otibazo Morimoto, 1961, Seleuca Pascoe, 1871 and Acallinus Morimoto, 1962 (all Curculionidae); species Oodes (Lachnocrepis) japonicus (Bates, 1873) (Carabidae: Licininae), Drusilla obliqua (Bernhauer, 1916) (Staphylinidae: Aleocharinae) and Coccotrypes advena Blandford, 1894 (Curculionidae: Scolytinae). The records of Anapleus Horn, 1873 (Histeridae) and Batraxis Reitter, 1882 (Staphylinidae: Pselaphinae) have been confirmed. The male of Sivacrypticus taiwanicus Kaszab, 1964 (Archeocrypticidae) is described for the first time. Gyrelon jenpani Hu, Fikáček & Matsumoto, sp. nov. (Cerylonidae) is described, illustrated, and compared with related species. DNA barcodes associated larvae of 42 species with adults, we are concisely illustrating some of these: Oodes japonicus, Perigona cf. nigriceps Dejean, 1831 (both Carabidae), Ptilodactyla sp. (Ptilodactylidae), Maltypus ryukyuanus Wittmer, 1970 (Cantharidae), Drusilla obliqua, Myrmecocephalus brevisulcus (Pace, 2008), Diochus sp., Mimopinophilus sp. (all Staphylinidae), Stelidota multiguttata Reitter, 1877, Lasiodites inaequalis (Grouvelle, 1914) (both Nitidulidae), Lagria scutellaris Pic, 1910, and Anaedus spinicornis Kaszab, 1973 (both Tenebrionidae). We also report the first cases of Rickettsia infections in Scydmaeninae and Pselaphinae. All data (sequences, metadata, and voucher photos) are made public in BOLD database and in a Zenodo Archive.

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Research Article Mon, 8 Jan 2024 15:53:39 +0200
Two new genera and four new species of Neotropical Eccritotarsini (Heteroptera, Miridae, Bryocorinae) https://dez.pensoft.net/article/104130/ Deutsche Entomologische Zeitschrift 71(1): 1-15

DOI: 10.3897/dez.71.104130

Authors: Eugenia Minghetti, Sara Itzel Montemayor, Pablo Matías Dellapé

Abstract: In this study two new genera and four new species of Eccritotarsini (Hemiptera: Heteroptera: Miridae: Bryocorinae) are described. Thomasomiris gen. nov. is established to accommodate Thomasomiris setosus sp. nov. (from Panama) and Egerocoris gen. nov., is described for E. ecuatorianus sp. nov. (the type species), E. dimorphus sp. nov. (both from Ecuador) and E. chaparensis sp. nov. (from Bolivia). Adult habitus and male genitalia photographs are provided for each species. A key to species is also provided and the affinities of the new genera are discussed.

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Research Article Mon, 8 Jan 2024 15:52:37 +0200
Two new species of Deuterophlebia Edwards, 1922 from Southwestern China (Diptera, Deuterophlebiidae) https://dez.pensoft.net/article/108096/ Deutsche Entomologische Zeitschrift 70(2): 387-401

DOI: 10.3897/dez.70.108096

Authors: Xuhongyi Zheng, Changfa Zhou

Abstract: The family Deuterophlebiidae is considered the most plesiomorphic Diptera family, with striking morphology and life history. In this study, we provide detailed descriptions and figures of two new species from Southwestern China: Deuterophlebia pseudopoda sp. nov. and Deuterophlebia pachychaeta sp. nov., along with genetic distances of COI sequences between the seven known Chinese species of Deuterophlebia Edwards, 1922. The two new species can be identified by the adults’ male terminalia, head structures, male and female antennae and mesothoracic spines or abdominal projections of the pupae. As a result, China is currently the country with the highest Deuterophlebia species diversity.

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Short Communication Mon, 23 Oct 2023 10:06:25 +0300
Comparative morphology of the larval mouthparts among six species of Notodontidae (Insecta, Lepidoptera), with discussions on their feeding habits and pupation sites https://dez.pensoft.net/article/107431/ Deutsche Entomologische Zeitschrift 70(2): 357-368

DOI: 10.3897/dez.70.107431

Authors: Jia-Xin Liu, Lu Jiang

Abstract: Larval mouthparts are significant organs for the individual development, morphologically related with feeding habits, and providing valuable characters for taxonomy and phylogenetic analysis. In previous studies, larval mouthparts revealed two identifying characters of Notodontidae. However, the evolutionary driving force and exact definition of these structures remain unsatisfactory. In this study, the larval mouthparts of Euhampsonia cristata (Butler, 1877), Fentonia ocypete (Bremer, 1861), Phalera assimilis (Bremer & Grey, 1853), Nerice davidi Oberthür, 1881, Cerura erminea (Esper, 1783) and Furcula furcula (Clerck, 1759) are morphologically observed and compared using scanning electron microscopy (SEM). The larval mouthparts of the six species are commonly equipped with paired maxillary sacs, congruent with the previous descriptions. However, the larval mouthparts of N. davidi are peculiar for bearing toothed mandibles, providing an exception of Notodontidae. Otherwise, the mouthparts exhibit morphological differences on mandibles, spinnerets, labral notches, and setal arrangements among the six species. The morphological diversity and the related feeding and pupation habits are briefly discussed.

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Research Article Fri, 29 Sep 2023 15:36:56 +0300
Phylogeny of genus Sichuana Shen & Yin, 2020 (Orthoptera, Tettigoniidae, Tettigoniinae) with four new species from Sichuan, China https://dez.pensoft.net/article/105870/ Deutsche Entomologische Zeitschrift 70(2): 337-355

DOI: 10.3897/dez.70.105870

Authors: Jun-Jie Gu, Chengjie Zheng, Su-Rong Jiang, Yanli Yue

Abstract: Four new species of Sichuana Shen & Yin, 2020 are described based on morphological comparison and molecular analysis: S. planicercata sp. nov., S. curvicercata sp. nov., S. longilamina sp. nov. and S. magnicerca sp. nov. Specimens showed some intraspecific variation of male tegmina and subgenital plates. The genes COI and 16S were used to analyze the genetic distance between species and COI was used to analyze the phylogenetic relationship of Sichuana.

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Research Article Fri, 29 Sep 2023 11:28:58 +0300
Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini) https://dez.pensoft.net/article/107425/ Deutsche Entomologische Zeitschrift 70(2): 311-335

DOI: 10.3897/dez.70.107425

Authors: Arnaud Faille, Sylvia Hofmann, Yeshitla Merene, David Hauth, Lars Opgenoorth, Yitbarek Woldehawariat, Joachim Schmidt

Abstract: The trechine beetle fauna (Coleoptera, Carabidae) of the Ethiopian Highlands is known to be highly diverse in species, and many species groups were recognized to be characterized by unusual character states of external and genital morphology. Earlier authors described several genera and subgenera of Ethiopian Trechina endemic to certain high mountains of the country. However, the relationships of these species groups and their evolutionary history are unknown so far. Here, we present the first molecular phylogenetic analysis of Ethiopian Trechina, detect several synonymic names under Trechus sensu lato, and introduce two new species groups to the country’s fauna: the monotypic genus Baehria Schmidt & Faille, gen. nov., with the type species B. separata sp. nov. from Mt. Choke in northern Ethiopia, and the Trechus subgenus Abunetrechus Schmidt & Faille, subgen. nov., with the type species T. bipartitus Raffray, 1885; this subgenus includes three species of northern Ethiopia. We show that the composition of the Ethiopian fauna is based on multiple events of immigration, which started simultaneously with or some million years after the Oligocene-Early Miocene orogenic events north and south of the Rift Valley. Our results support the habitat island hypothesis for the evolution of the Ethiopian highland fauna. We found no evidence for an alternative hypothesis assuming a close connection of the Trechina immigration to Ethiopia and Pleistocene cooling. We, thus, conclude that the geomorphological development rather than the climatic changes are the main drivers of the diversification of the high-altitude Trechina fauna in Ethiopia.

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Research Article Wed, 27 Sep 2023 17:44:12 +0300
A morphology-based revision and phylogenetic analysis of the Pterostichus macrogenys species group (Coleoptera, Carabidae) and implications for differentiation of the species group https://dez.pensoft.net/article/107322/ Deutsche Entomologische Zeitschrift 70(2): 291-310

DOI: 10.3897/dez.70.107322

Authors: Kôji Sasakawa, Yoshiji Mitsuduka

Abstract: The Pterostichus macrogenys species group is an endemic subterranean Japanese carabid clade that provides intriguing material for studying morphological differentiation, speciation and interspecific relationships. However, its diversity remains not fully explored. We investigated specimens from northern Tohoku District, an area where knowledge of this species group is notably limited. Our research led to the description of three new species: P. namahage sp. nov., P. kamurosanus sp. nov. and P. atsumidakensis sp. nov. We also updated distribution records for three known species: P. asahinus Habu & Baba, 1960; P. kitakamisanus Sasakawa, 2005; and P. chokaisanus Sasakawa, 2009. This report includes both the expanded distribution ranges of the known species and the discovery of a new sympatric species pair (P. chokaisanus and P. asahinus). We conducted a morphological phylogenetic analysis of all but one species, for which no male specimens were available, accounting for a total of 42 species within the group. The resulting phylogenetic tree implies that the initial differentiation of this species group originated on the Sea of Japan side, in the northern part of their current distribution, followed by dispersion to other areas and subsequent differentiation. Additionally, our findings indicate that sympatric species of varying body sizes are distantly related phylogenetically. These insights into the differentiation process align with regional distribution patterns of species-level diversity and sympatric sites.

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Research Article Thu, 17 Aug 2023 18:01:47 +0300
Taxonomic review of the genus Sinopanorpa Cai & Hua, 2008 (Mecoptera, Panorpidae) with descriptions of two new species https://dez.pensoft.net/article/104621/ Deutsche Entomologische Zeitschrift 70(2): 283-290

DOI: 10.3897/dez.70.104621

Authors: Yuan Hua, Kai Gao, Lianxi Xing

Abstract: The scorpionfly genus Sinopanorpa Cai & Hua, 2008 is endemic to the mountain regions of central China, currently consisting of four species: Sinopanorpa tincta (Navás, 1931), S. digitiformis Huang & Hua, 2008, S. nangongshana Cai & Hua, 2008, and S. baokangensis Wang, 2021. Here, the genus is taxonomically reviewed, with descriptions of two new species: Sinopanorpa minshanicola sp. nov. from the Minshan Mountains and Sinopanorpa shennongjiaica sp. nov. from the Shennongjia Mountains, increasing the species number of the genus to six. An updated key to species of Sinopanorpa is provided.

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Research Article Thu, 27 Jul 2023 16:20:36 +0300
An “American” silkmoth endemic to Himalayas, part I: life history and natural distribution of Antheraea compta Rothschild, 1899 (Lepidoptera, Saturniidae) https://dez.pensoft.net/article/102952/ Deutsche Entomologische Zeitschrift 70(2): 261-282

DOI: 10.3897/dez.70.102952

Authors: Zhengyang Liu

Abstract: Females of the uncommon Antheraea compta Rothschild, 1899 were collected at the natural habitat in the Tibetan Sub-Himalayas, descendants of which were reared on Quercus yunnanensis (Fagaceae) successfully, with all the preimaginal instars recorded morphologically. Correlated characters revealed a close relationship between this taxon and New World Antheraea spp., suggesting more attention and protection towards this key species is necessary in the future.

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Research Article Mon, 24 Jul 2023 13:01:56 +0300
The snakeflies of the Mediterranean islands: review and biogeographical analysis (Neuropterida, Raphidioptera) https://dez.pensoft.net/article/101559/ Deutsche Entomologische Zeitschrift 70(1): 175-218

DOI: 10.3897/dez.70.101559

Authors: Horst Aspöck, Ulrike Aspöck

Abstract: The Mediterranean region representsone of the hotspots of biodiversity of many organisms and among these also of Raphidioptera. With about 250 known species world-wide Raphidioptera is the smallest order of extant holometabolous insects. The Mediterranean region harbors 113 species (= >45% of the world fauna), and of these 33 (= >13% of the world fauna) have been found on islands of the Mediterranean Sea. In the course of the past 50 years most of the larger islands have been intensively explored for Raphidioptera, particularly all islands (except the Baleares) which had no connection to the mainland throughout the whole Pleistocene. Altogether, 11 species of Raphidioptera (9 Raphidiidae, 2 Inocelliidae) are endemic to (usually)one, rarely to a few Mediterranean islands: Phaeostigma (Ph.) euboica (Euboea), Ph. (Aegeoraphidia) prophetica (Rhodes), Ph. (Ae.) karpathana (Karpathos), Ph. (Ae.) biroi (Crete), Ph. (Superboraphidia) minois (Crete), Subilla principiae (Sardinia), S. colossea (Rhodes), Raphidia (R.) peterressli (Chios), R. (R.) ariadne (Crete), Fibla (F.) maclachlani (Corsica, Sardinia, Sicily), F. (Reisserella) pasiphae (Crete). In the Aegean Sea there is a remarkable difference between the eastern and the western islands marked by the so-called Rechinger line based on results of the analysis of the flora. The Raphidioptera confirm this line impressively. It is rather unlikely that further Raphidioptera species endemic to Mediterranean islands are still to be detected – with one exception: the Baleares. A few additional species known from various parts of the continents surrounding the Mediterranean Sea will probably be found on islands so that a total number may be around 40, possibly around 45 species. The possible ways and times of colonization of the islands by Raphidioptera are discussed. The present paper may serve as a basis for studies on natural and particularly on anthropogenic dispersals of snakeflies from island to island, from mainland to islands, and from islands to the mainland in connection with phylogeographic investigations.

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Research Article Wed, 3 May 2023 15:07:38 +0300
Resolving inconsistencies between Plötz’s descriptions and presumed type specimens of some Hesperiidae (Lepidoptera) https://dez.pensoft.net/article/98280/ Deutsche Entomologische Zeitschrift 70(1): 159-174

DOI: 10.3897/dez.70.98280

Authors: Jing Zhang, Qian Cong, Leina Song, Jinhui Shen, Théo Léger, Gerardo Lamas, Olaf H. H. Mielke, Nick V. Grishin

Abstract: Comparing specimens curated in MfN as primary types of Hesperiidae names proposed by Carl Plötz with the original descriptions and unpublished drawings reveals a number of inconsistencies that we address. Lectotypes are designated for Telegonus labriaris Butler, 1877, Eudamus jalapus Plötz, 1881, and Apaustus interpunctata Plötz, 1884. Neotypes are designated for Netrocoryne seneca Plötz, 1882 and Hesperia irma Plötz, 1882. Hesperia ulphila Plötz, 1883 is treated as a nomen dubium. As a result of these designations, the following are junior objective synonyms: Netrocoryne seneca Plötz, 1882 of Telegonus labriaris Butler, 1877 and Hesperia irma Plötz, 1882 of Pamphila irma Möschler, 1879, the latter two names being homonyms. In all these instances, the original descriptions are satisfied, and, except for A. interpunctata, the current application of these names is unaffected. Taxonomically, Callimormus diaeses Schaus, 1902, stat. rest. is a valid species, not a synonym of A. interpunctata, which is a junior subjective synonym of Eutocus vetulus (Mabille, 1883).

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Research Article Mon, 3 Apr 2023 15:39:55 +0300
The genus Anacaena Thomson from the Ryukyu Archipelago of Japan (Coleoptera, Hydrophilidae) https://dez.pensoft.net/article/96994/ Deutsche Entomologische Zeitschrift 70(1): 143-157

DOI: 10.3897/dez.70.96994

Authors: Yûsuke N. Minoshima, Yuuki Kamite, Martin Fikáček

Abstract: We review the genus Anacaena Thomson, 1859 from the Ryukyu Archipelago, southern Japan. Three aquatic species are recognised: A. torikaii sp. nov. from Amami-ôshima Island, A. okinawana sp. nov. from Okinawa-jima Island and Kerama Islands, and A. kumejimana sp. nov. from Kumejima Island. All three species are very similar, with the morphology of the aedeagus being essential for a reliable identification. Dorsal colouration is also useful as a diagnostic character, despite some variation within species. We observe a possible geography-based variation between A. okinawana from Okinawa-jima I. and the neighbouring Kerama Is., but we treat both populations as conspecific based on genital morphology. Anacaena kumejimana and A. okinawana share many morphological characters possibly indicating their close relationship. We compare the endemism of aquatic Hydrophilidae in the Ryukyu Archipelago to that in other groups of aquatic beetles: the proportion of endemic species is higher in aquatic Hydrophilidae than in Dytiscidae, but much lower than in stream-inhabiting Hydraenidae and Elmidae. A list of Japanese species of Anacaena and a key to the Japanese species of the genus are provided.

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Research Article Fri, 24 Mar 2023 10:45:37 +0200
Caribbean pygmy jumping leaves (Tetrigidae, Cladonotinae, Choriphyllini) https://dez.pensoft.net/article/98982/ Deutsche Entomologische Zeitschrift 70(1): 129-141

DOI: 10.3897/dez.70.98982

Authors: Josip Skejo, Sheyla Yong, Domagoj Bogić, Niko Kasalo

Abstract: The tribe Choriphyllini Cadena-Castañeda & Silva, 2019 consists of only two genera, Choriphyllum Serville, 1838 and Phyllotettix Hancock, 1902b, combining leaf-like Caribbean wide-nosed pygmy grasshoppers (Tetrigidae, Cladonotinae). The genus Choriphyllum has four species, C. bahamense Perez-Gelabert & Otte, 1999 from the Bahamas (Hummingbird Cay Island), and C. sagrai Serville, 1838, C. saussurei Bolívar, 1887 and C. wallaceum Skejo, Kasalo & Yong, sp. nov. from Cuba. The gender of C. bahamensis is changed to C. bahamense in order to match the grammatical gender of the genus, which is neuter. Silva’s designation of C. sagrai, the type species of Choriphyllum, as nomen dubium in 2019 was incorrect as Serville’s drawing clearly points to this species, endemic to Cuba, so nothing about its identity is doubtful. The genus Phyllotettix is endemic to Jamaica, where four species live: P. rhombeus (Felton, 1765), P. foliatus (Hancock, 1902a), P. compressus (Thunberg, 1815) and P. plagiatus (Walker, 1871), comb. nov. of Choriphyllum plagiatum. Choriphyllum westwoodi Hancock, 1902a, syn. nov. is synonymous with Phyllotettix rhombeus and not with P. compressus. A new subgenus is established, Phyllotettix (Rhombotettix) subgen. nov. for P. (R.) plagiatus comb. nov. and P. (R.) rhombeus. A new species complex, Phyllotettix (compressus) sp. complex nov. is established for two morphologically close species, P. compressus, and P. foliatus; while another new species complex, Choriphyllum (sagrai) sp. complex nov. includes C. sagrai and C. wallaceum sp. nov. An annotated key to genera, subgenera, species groups and species is provided.

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Research Article Fri, 24 Mar 2023 10:44:03 +0200
Revisionary notes on Feuerborniella Vaillant, 1971, with the first record of the genus from the Afrotropical region (Diptera, Psychodidae) https://dez.pensoft.net/article/97465/ Deutsche Entomologische Zeitschrift 70(1): 121-127

DOI: 10.3897/dez.70.97465

Authors: Gunnar Mikalsen Kvifte, Santiago Jaume-Schinkel

Abstract: We establish a diagnosis for Feuerborniella Vaillant, 1971, based on a re-description of its type species, Feuerborniella obscura (Tonnoir, 1919) and comment on earlier diagnoses. Feuerborniella sinefurcata Kvifte & Jaume-Schinkel, sp. nov. is described, based on material from Tanzania, representing the second Afrotropical species of the genus following Psychoda morogorica Wagner & Andersen, 2007 which we treat as Feuerborniella morogorica comb. nov. We furthermore review earlier combinations, transferring Philosepedon ensiger Quate, 1996 and Philosepedon longistylus Quate, 1996 to Feuerborniella comb. nov., and briefly discuss generic limits with Quatiella Botosaneanu & Vaillant, 1970 and Nielseniella Vaillant, 1971.

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Research Article Thu, 16 Mar 2023 18:32:48 +0200
A remarkable new genus of Nevrorthidae (Neuroptera, Osmyloidea) from mid-Cretaceous Kachin amber of northern Myanmar https://dez.pensoft.net/article/98873/ Deutsche Entomologische Zeitschrift 70(1): 113-120

DOI: 10.3897/dez.70.98873

Authors: Hiroshi Nakamine, Shûhei Yamamoto, Yui Takahashi, Xingyue Liu

Abstract: A remarkable new genus and species of Nevrorthidae, Sisyroneurorthus aspoeckorum gen. et sp. nov., is described from mid-Cretaceous Kachin amber of northern Myanmar. This new species is the second Mesozoic representative of Nevrorthidae. The distal fusion of ScP and RA in the forewing of this new genus is shared by most Sisyridae. Given the probable sister-group relationship between Nevrorthidae and Sisyridae, our study sheds light on the evolution and morphological diversity of Nevrorthidae in the Mesozoic era.

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Research Article Thu, 16 Mar 2023 18:30:20 +0200
Revision of the genus Eotrechus Kirkaldy (Hemiptera, Heteroptera, Gerridae), with descriptions of six new species https://dez.pensoft.net/article/97117/ Deutsche Entomologische Zeitschrift 70(1): 69-111

DOI: 10.3897/dez.70.97117

Authors: Anh Duc Tran, Herbert Zettel, Robert W. Sites

Abstract: Species of the Asian genus Eotrechus Kirkaldy, 1902 (Hemiptera, Heteroptera, Gerromorpha, Gerridae, Eotrechinae) are atypical water striders because they live on the near-vertical rock surfaces of waterfalls. The taxonomy of the genus is revised and six new species are described: E. steineri sp. nov., E. kerberos sp. nov., E. thai sp. nov., E. konkakinh sp. nov., E. boukali sp. nov., and E. anderseni sp. nov. Supplementary descriptions of the apterous female and macropterous male of E. fuscus Basu, Chandra & Venkatesan, 2017 are provided. Additional and updated distribution data of species of Eotrechus are reported. Eotrechus fuscus is reported from Myanmar for the first time; E. brevipes is reported from Nepal for the first time; E. hygropetricus Andersen, 1982 is reported from southern Thailand for the first time, which extends the southernmost limit of the known distribution of the genus. A revised key to the species of Eotrechus is also provided.

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Research Article Thu, 16 Mar 2023 18:28:40 +0200
Mouthpart palp sensilla of basal Trichoptera families https://dez.pensoft.net/article/98752/ Deutsche Entomologische Zeitschrift 70(1): 55-68

DOI: 10.3897/dez.70.98752

Authors: Kseniia T. Abu Diiak, Vladimir D. Ivanov, Stanislav I. Melnitsky, Mikhail Yu. Valuyskiy, Alexandra A. Puyto

Abstract: A comparative SEM study of palp sensory surfaces in 25 caddisfly species representing seven families reveals seven types of sensilla: long trichoid, blunt chaetoid, campaniform, mushroom-like pseudoplacoid, petaloid, thick basiconic and thin basiconic sensilla. Long trichoid and chaetoid sensilla are present on all segments of both pairs of palps. First and second segments of maxillary palps bear groups of long and sclerotised chaetoid sensilla on their medial surface. Other segments of maxillary palps and all segments of labial palps have shorter and thinner chaetoid sensilla mainly on their ventromedial surfaces. Campaniform sensilla usually occur on the first segment of labial palps and second segment of maxillary palps. Mushroom-like pseudoplacoid sensilla may occupy all palp segments or only distal ones. Petaloid sensilla form sensory fields on apical segments of both pairs of palps in most studied species. Thick basiconic sensilla occur only in apical sensory complexes on tips of maxillary and labial palps. A comparison with the Lepidoptera suggests the similarity in palp sensilla and conservative evolution of the palp surface. The reconstructed ground plan for the palp sensory surfaces in Trichoptera and Amphiesmenoptera is provided.

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Research Article Thu, 16 Mar 2023 18:25:17 +0200
Revision of the South African leafhopper genus Hadroca Theron (Hemiptera, Auchenorrhyncha, Cicadellidae, Bonaspeiini) https://dez.pensoft.net/article/90851/ Deutsche Entomologische Zeitschrift 70(1): 13-54

DOI: 10.3897/dez.70.90851

Authors: Michael Stiller

Abstract: The leafhopper genus Hadroca Theron, 1974 with type species Euscelis ramosa Naudé, 1926 is re-defined here by examination of long series of specimens with varied color and wing polymorphic males and females. Five new species are added: Hadroca alacaudella sp. nov., H. alavittata sp. nov., H. bualacauda sp. nov., H. hapsistylis sp. nov. and H. treichroa sp. nov. The genus is known only from South Africa, mostly from the Fynbos Biome. All species have the aedeagus C-shaped and uniformly sclerotized, the shaft gradually tapered, commonly edentate, rarely with single, subapical, anterior process and the style with distal apophysis curved ventrad. A key to species is provided, with discussion on the differentiation from similar genera such as Bloemia Theron, 1974, Basutoia Linnavuori, 1961 and Tzitzikamaia Linnavuori, 1961 and distribution maps.

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Research Article Fri, 3 Feb 2023 17:48:41 +0200
Four new species of Ischnodemus (Hemiptera, Heteroptera, Blissidae) and additional records from Argentina https://dez.pensoft.net/article/94683/ Deutsche Entomologische Zeitschrift 69(2): 283-295

DOI: 10.3897/dez.69.94683

Authors: Pablo M. Dellapé, María C. Melo

Abstract: Four new species of the genus Ischnodemus Fieber, 1837 (Hemiptera, Heteroptera, Blissidae) are described from Argentina: I. nigromaculatoides sp. nov., I. formosus sp. nov., I. infernalis sp. nov. and I. correntinus sp. nov. The known distribution is extended for three previously recorded species in the country. The species Ischnodemus brusai Dellapé & Montemayor, 2009 from Peru is transferred to Patritiodemus Slater & Ahmad, 1971. Dorsal habitus photographs and detailed images of the pygophore and right paramere of the new species and a key to the Argentinean species of Ischnodemus are given.

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Research Article Thu, 15 Dec 2022 10:35:04 +0200
Integrative taxonomy of the Lauraceae-feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan https://dez.pensoft.net/article/89864/ Deutsche Entomologische Zeitschrift 69(2): 219-281

DOI: 10.3897/dez.69.89864

Authors: Jun Souma

Abstract: Stephanitis Stål, 1873 (Hemiptera, Heteroptera, Tingidae, Tinginae, Tingini) is one of the most diverse lace bug genera in East Asia with over 50 species, many of them feeding on Ericaceae and Lauraceae. The Japanese species feeding on Lauraceae have high intraspecific and poor interspecific variation, so identification of some previous distribution records (mainly S. (Norba) aperta Horváth, 1912 and closely related taxa) is doubtful. In the present study, 5,080 specimens of Lauraceae-feeding species of Stephanitis from Japan were identified to species based on morphology, and DNA barcodes of the mitochondrial cytochrome c oxidase subunit (COI) region for 53 of those individuals were also obtained. The combined morphological and molecular evidence dataset showed that 10 species of Stephanitis feed on lauraceous trees in Japan: S. (Stephanitis) ambigua Horváth, 1912, S. (Norba) aperta, S. (N.) exigua Horváth, 1912, S. (N.) hayashii sp. nov., S. (N.) hiurai Takeya, 1963, S. (N.) ishikawai sp. nov., S. (N.) mendica Horváth, 1912, S. (S.) tabidula Horváth, 1912, S. (S.) takeyai Drake & Maa, 1955 and S. (S.) tomokunii sp. nov. Additionally, a new synonymy, S. (S.) fasciicarina Takeya, 1931, syn. nov. with S. (S.) tabidula, is proposed, a key for identifying the 10 species is provided and their distribution ranges and host plant relationships are presented.

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Research Article Thu, 15 Dec 2022 10:34:53 +0200
Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta) https://dez.pensoft.net/article/84080/ Deutsche Entomologische Zeitschrift 69(2): 151-218

DOI: 10.3897/dez.69.84080

Authors: Andrew Liston, Marko Mutanen, Mikk Heidemaa, Stephan M. Blank, Niina Kiljunen, Andreas Taeger, Matti Viitasaari, Veli Vikberg, Saskia Wutke, Marko Prous

Abstract: While working on an identification guide to the sawflies of Fennoscandia, we encountered numerous taxonomic problems, for some of which we present solutions. Dicrostema Benson, 1952 is a new synonym of Phymatoceropsis Rohwer, 1916, and not congeneric with Paracharactus MacGillivray, 1908. Two species occurring in Europe are transferred to Phymatoceropsis. Dolerus aericepsellus Heidemaa and Mutanen sp. nov. and Heptamelus viitasaarii Liston, Mutanen and Prous sp. nov. are described from Finland. Abia brevicornis Leach, 1817 nom. rev. is the valid name of Abia nitens auct. nec Linnaeus, and Abia nitens (Linnaeus, 1758) is the valid name for what has recently been called Abia sericea (Linnaeus, 1767). Tenthredo haemorrhoidalis Fabricius, 1781 is treated as an unplaced species of Hymenoptera, possibly Ichneumonoidea. Calameuta variabilis (Mocsáry, 1886) is the valid name of the species recently generally called C. haemorrhoidalis. Claremontia confusa (Konow, 1886) sp. rev. and Claremontia brevicornis (Brischke, 1883) are distinct species. Dolerus coracinus (Klug, 1818) is the valid name for D. anthracinus auct. Dolerus anthracinus (Klug, 1818) is a valid species similar to D. nitens Zaddach, 1859. Dolerus coruscans Konow, 1890 sp. rev. is a valid species. Dolerus junci (Stephens, 1835) is the valid name for Dolerus cothurnatus auct. Dolerus timidus (Klug, 1818) sp. rev. is distinguished from the similar D. pratensis (Linnaeus, 1758). A neotype is designated for Astatus punctatus Klug, 1803. Lectotypes are designated for 39 nominal species. 29 species group names are new junior synonyms. We present data on some species recently collected for the first time in Finland, including first records for the Palaearctic and West Palaearctic.

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Research Article Tue, 26 Jul 2022 21:08:45 +0300
A redefinition of Umbrageocoris with new species and new combinations (Heteroptera, Lygaeoidea, Geocoridae) https://dez.pensoft.net/article/85584/ Deutsche Entomologische Zeitschrift 69(2): 139-150

DOI: 10.3897/dez.69.85584

Authors: Péter Kóbor

Abstract: The geocorine (Hemiptera: Heteroptera: Lygaeoidea: Geocoridae) true bug genus Umbrageocoris Kóbor, 2019 is redefined based on new morphological information from newly acquired specimens. Two new species are described: U. boonei sp. nov. from continental Indomalaya and U. malipatili sp. nov. from Australia; two new combinations are proposed: U. elegantulus (Distant, 1904), comb. nov., and U. woodwardi (Malipatil, 1994), comb. nov. (both transferred from Geocoris Fallén, 1814). Keys, diagnoses, and distribution data to the discussed species are provided. Hypotheses on the origin of Umbrageocoris and its relationship to other geocorine genera in the region are formulated. New country records: U. elegantulus (Papua New Guinea), U. maai maai (Thailand, Laos) and U. woodwardi (Papua New Guinea).

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Research Article Thu, 21 Jul 2022 21:04:04 +0300
Review of the genus Sericocampsomeris Betrem, 1941 (Hymenoptera, Scoliidae) from China https://dez.pensoft.net/article/83335/ Deutsche Entomologische Zeitschrift 69(2): 125-138

DOI: 10.3897/dez.69.83335

Authors: Hua-Yan Chen, Zhen Liu, Zheng Wang, Sheng-Jie Yang, Shi-Xiao Luo

Abstract: The Chinese species of the genus Sericocampsomeris Betrem, 1941 are revised, based on morphology. Four species are recognised, of which one species is described as new, i.e. Sericocampsomeris punctata Liu & Chen, sp. nov., for which we also provide molecular sequence data to supplement its identity. A key to species of Sericocampsomeris from China and a distribution map are provided. Furthermore, we identify Sericocampsomeris punctata Liu & Chen, sp. nov. as a pollinator of Elaeocarpus sp. (Elaeocarpaceae) and recognise Leucopholis rorida (Fabricius, 1801) as a host of Sericocampsomeris rubromaculata (Smith, 1855).

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Research Article Thu, 21 Jul 2022 21:03:15 +0300
Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini) https://dez.pensoft.net/article/83840/ Deutsche Entomologische Zeitschrift 69(1): 71-122

DOI: 10.3897/dez.69.83840

Authors: Borislav Guéorguiev, David W. Wrase, Thorsten Assmann, Jan Muilwijk, Patrice Machard

Abstract: Species of the subgenus Eremoderus Jeanne, 1996, genus Platyderus Stephens, 1827, occurring in continental Africa (excluding Macaronesia) and southwest Asia, are taxonomically revised. The following new species groups and species are defined and described, “weiratheri” group: Platyderus (Eremoderus) chatzakiae, sp. nov. (type locality: Greece, Kalymnos Island, near Stimenia Village); “iranicus-vanensis” group: Platyderus (Eremoderus) felixi, sp. nov. (type locality: Iran, Chahar Mahal va Bakhtiari Province, 10 km west of Naghan Town); Platyderus (Eremoderus) iranicus, sp. nov. (type locality: Iran, Chahar Mahal va Bakhtiari Province, 7 km NE Naghan Town); Platyderus (Eremoderus) vanensis, sp. nov. (type locality: Turkey, Van Province, Gevaş Town); Platyderus (Eremoderus) vrabeci, sp. nov. (type locality: Turkey, Nemrut Daği); “lassallei” group: Platyderus (Eremoderus) lassallei, sp. nov. (type locality: Iran, Mazandaran Province, between Nur City and Lavij Village); “davatchii” group: Platyderus (Eremoderus) klapperichi, sp. nov. (type locality: Iran, Mazandaran Province, Damavand, 2000 m); “afghanistanicus” group: Platyderus (Eremoderus) afghanistanicus, sp. nov. (type locality: Afghanistan, “Habatah”); “languidus” group: Platyderus (Eremoderus) arabicus, sp. nov. (type locality: Saudi Arabia, “Hedjaz”); Platyderus (Eremoderus) brunki, sp. nov. (type locality: Republic of Yemen, Thula District, between Kaukaban and Shibam); Platyderus (Eremoderus) irakensis, sp. nov. (type locality: Iraq, Ar Rutba District, 115 km E Ar-Rutbah Town); Platyderus (Eremoderus) jordanensis, sp. nov. (type locality: Jordan, Al-Betrā’ District, Little Petra). Six previously described species — P. brunneus Karsch, P. insignitus Bedel, P. languidus Reiche & Saulcy, P. ledouxi Morvan, P. taghizadehi Morvan, and P. weiratheri Mařan — are redescribed based on type and/or non-type material. P. davatchii Morvan placed as a member of the subgenus was not treated due to the lack of material available for study. The following new nomenclature acts are proposed: Platyderus brunneus Karsch, 1881, stat. rev., is removed from synonymy with Feronia languida Reiche & Saulcy, 1855; Platyderus elegans Bedel, 1900, syn. nov., is proposed as junior synonym of Platyderus brunneus Karsch, 1881; Platyderus ferrantei Reitter, 1909 is proposed as subspecies Platyderus brunneus ferrantei Reitter, 1909, stat. nov. In order to preserve the stability of nomenclature, lectotypes are designated for: Feronia languida Reiche & Saulcy, Platyderus brunneus Karsch, and Platyderus weiratheri Mařan. Keys to identification of the male and female specimens of the species from the regions studied are provided.

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Research Article Mon, 13 Jun 2022 17:04:17 +0300
Identity of Zorotypus juninensis Engel, 2000, syn. nov. revealed: it is conspecific with Centrozoros hamiltoni (New, 1978) (Zoraptera, Spiralizoridae) https://dez.pensoft.net/article/83154/ Deutsche Entomologische Zeitschrift 69(1): 65-70

DOI: 10.3897/dez.69.83154

Authors: Petr Kočárek, Ivona Horká

Abstract: Zorotypus juninensis Engel, 2000, was previously diagnosed based on the external morphology of female and male specimens without description of the male copulatory organ, which is an important character for classification in Zoraptera. Based on a detailed morphological study of the Zorotypus juninensis Engel, 2000 type collections deposited in the American Museum of Natural History in New York, and based on the comparison with the holotype male of Centrozoros hamiltoni (New, 1978), we have determined that these two species are conspecific. We therefore formally synonymize Zorotypus juninensis Engel, 2000, syn. nov. with Centrozoros hamiltoni (New, 1978). Morphological characters and phylogenetic relationships of Centrozoros Kukalova-Peck & Peck, 1993 are also discussed in this report.

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Research Article Wed, 8 Jun 2022 10:41:03 +0300
Revision of the genus Woldstedtius Carlson, 1979 (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan https://dez.pensoft.net/article/80492/ Deutsche Entomologische Zeitschrift 69(1): 45-64

DOI: 10.3897/dez.69.80492

Authors: Shunsuke Morishita, Kyohei Watanabe

Abstract: Japanese species of the genus Woldstedtius Carlson, 1979 are revised. Nine species are recorded from Japan, including two new species, W. alpicola sp. nov. and W. punctatus sp. nov. Woldstedtius biguttatus (Gravenhorst, 1829) is newly recorded from Japan. Taxonomic status of W. flavolineatus kuroashii (Uchida, 1957) is changed from the subspecies of W. flavolineatus (Gravenhorst, 1829) to a separated species. Woldstedtius holarcticus (Diller, 1969) is newly synonymized under W. kuroashii (Uchida, 1957). A key to Japanese species of this genus is provided.

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Review Article Mon, 4 Apr 2022 14:20:05 +0300
Character analysis and descriptions of Eocene sphodrine fossils (Coleoptera, Carabidae) using light microscopy, micro-CT scanning, and 3D imaging https://dez.pensoft.net/article/79931/ Deutsche Entomologische Zeitschrift 69(1): 19-44

DOI: 10.3897/dez.69.79931

Authors: Joachim Schmidt, Stephan Scholz, Kipling Will

Abstract: Of the 12 specimens of Calathus-like sphodrine beetles presently known from Baltic and Rovno amber deposits, 11 specimens were investigated using light microscopy, micro-CT scanning, and 3D imaging techniques. For the first time, many significant diagnostic characters of the external morphology and male and female genitalia of Eocene Sphodrini were studied in detail. Based on these data, three fossil species are diagnosed and placed in a natural group characterized by a derived pattern in elytral chaetotaxy and microsculpture and therefore the genus Quasicalathus Schmidt & Will, gen. nov. is described to comprise these species. Due to the presence of a styloid right paramere, Quasicalathus gen. nov. is considered a member of the sphodrine “P clade” of Ruiz et al. (2009). However, given the absence of synapomorphies of any species group of the P clade, the systematic position of Quasicalathus gen. nov. within this clade remains unresolved. The Baltic amber species Calathus elpis Ortuño & Arillo, 2009 is redescribed based on additional, fossil, non-holotype material and transferred to Quasicalathus gen. nov. Identification of the additional C. elpis fossil material remains slightly uncertain due to the non-availability of the holotype for direct comparison coupled with doubts regarding the accuracy of certain character states presented in its original description. Two species are newly described: Quasicalathus agonicollis Schmidt & Will, sp. nov., from Baltic amber, and Q. conservans Schmidt & Will, sp. nov., from Rovno amber.

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Research Article Wed, 9 Feb 2022 11:11:21 +0200
Phylogeny and age of cockroaches: a reanalysis of mitogenomes with selective fossil calibrations https://dez.pensoft.net/article/68373/ Deutsche Entomologische Zeitschrift 69(1): 1-18

DOI: 10.3897/dez.69.68373

Authors: Xin-Ran Li

Abstract: In spite of big data and new techniques, the phylogeny and timing of cockroaches remain in dispute. Apart from sequencing more species, an alternative way to improve the phylogenetic inference and time estimation is to improve the quality of data, calibrations and analytical procedure. This study emphasizes the completeness of data, the reliability of genes (judged via alignment ambiguity and substitution saturation), and the justification for fossil calibrations. Based on published mitochondrial genomes, the Bayesian phylogeny of cockroaches and termites is recovered as: Corydiinae + (((Cryptocercidae + Isoptera) + ((Anaplectidae + Lamproblattidae) + (Tryonicidae + Blattidae))) + (Pseudophyllodromiinae + (Ectobiinae + (Blattellinae + Blaberidae)))). With two fossil calibrations, namely, Valditermes brenanae and Piniblattella yixianensis, this study dates the crown Dictyoptera to early Jurassic, and crown Blattodea to middle Jurassic. Using the ambiguous ‘roachoid’ fossils to calibrate Dictyoptera+sister pushes these times back to Permian and Triassic. This study also shows that appropriate fossil calibrations are rarer than considered in previous studies.

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Research Article Fri, 7 Jan 2022 13:24:28 +0200
New species of Omma Newman from mid-Cretaceous Burmese amber (Coleoptera, Archostemata, Ommatidae) https://dez.pensoft.net/article/74174/ Deutsche Entomologische Zeitschrift 68(2): 341-348

DOI: 10.3897/dez.68.74174

Authors: Yan-Da Li, Di-Ying Huang, Chen‑Yang Cai

Abstract: A new fossil species of the extant archostematan genus Omma Newman, O. forte sp. nov., is reported from mid-Cretaceous amber from northern Myanmar. The extinct ommatid genus, Cionocups Kirejtshuk from the same deposit, is considered as a junior synonym of Omma, and C. manukyani is transferred to Omma, as O. manukyani comb. nov. A key to species of Omma from Burmese amber is also provided.

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Research Article Tue, 30 Nov 2021 12:23:58 +0200
Limnomma, a new genus of Ommatidae from the Middle Jurassic Daohugou beds (Coleoptera, Archostemata) https://dez.pensoft.net/article/71880/ Deutsche Entomologische Zeitschrift 68(2): 299-308

DOI: 10.3897/dez.68.71880

Authors: Yan-Da Li, Erik Tihelka, Hong Pang, Di-Ying Huang, Chen‑Yang Cai

Abstract: The relictual archostematan beetle family Ommatidae attained high diversity during the Mesozoic. Despite their once high taxonomic diversity and morphological disparity, many Mesozoic ommatid taxa remain poorly understood, partly due to limited preservation. Here we report an exceptionally well-preserved fossil, which we describe as a new ommatid genus and species, Limnomma daohugouense gen. et sp. nov., from the mid-Jurassic Daohugou Lagerstätte in Northeast China. The new genus can be most easily distinguished from other ommatids by the presence of a circular non-tuberculate region on ventrite 5. The new taxon is discussed in relation to the classification of the Mesozoic genera Brochocoleus and Burmocoleus.

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Research Article Fri, 3 Sep 2021 17:46:45 +0300
Revision of the genus Xanthopimpla Saussure, 1892 (Hymenoptera, Ichneumonidae, Pimplinae) from Japan https://dez.pensoft.net/article/69768/ Deutsche Entomologische Zeitschrift 68(2): 269-297

DOI: 10.3897/dez.68.69768

Authors: Kyohei Watanabe, Rikio Matsumoto

Abstract: Japanese species of the genus Xanthopimpla Saussure, 1892 are revised. A total of 15 species are recorded from Japan, including three new species, X. nipponensis sp. nov. X. sylvicola sp. nov. and X. yoshimurai sp. nov. and three species, X. honorata (Cameron, 1899), X. minuta Cameron, 1905 and X. trias Townes & Chiu, 1970, newly recorded from Japan. Host, habitat, overwintering and distribution patterns of Japanese Xanthopimpla species are discussed.

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Research Article Wed, 1 Sep 2021 17:46:05 +0300
Where did the Central European populations of Ornatoraphidia flavilabris (Costa) come from? (Neuropterida, Raphidioptera, Raphidiidae) https://dez.pensoft.net/article/70814/ Deutsche Entomologische Zeitschrift 68(2): 249-259

DOI: 10.3897/dez.68.70814

Authors: Horst Aspöck, Ulrike Aspöck, Julia Walochnik, Edwin Kniha

Abstract: Ornatoraphidia flavilabris (Costa, 1851) is one of 15 snakefly species occurring in southern parts of Central Europe. It is a polycentric Mediterranean faunal element with refugia in the Apennine Peninsula and the Balkan Peninsula. Two phylogeographic questions are dealt with in this paper: (1) Is it possible to differentiate, morphologically or genetically, the Balkanic populations from the Italian? (2) Did the species reach Central Europe from the Balkan or Apennine Peninsula? These questions were investigated using morphological and molecular biological methods. No morphological characters were uncovered which could serve to differentiate specimens from either distribution center. However, differences were detected in cox1, cox3 and 28S genes which allow for a reliable differentiation. Central European populations were largely identical with populations from Italy, but distinctly different from specimens from Greece. This could lead one to assume that the species migrated from Italy to Central Europe, although colonization from the southeast would appear easier due to more favorable orographic conditions. This discrepancy may be explained by the apparent absence of O. flavilabris from the large central part of the Balkan Peninsula, so that a gap exists between the southern and northern areas inhabited by O. flavilabris. Moreover, the species does not occur in eastern parts of Europe. Thus it would be more probable to assume that the occurrence of the species in the northwest Balkan Peninsula can be traced to migrations from the Apennine Peninsula to areas north and northeast of the Adriatic Sea, where O. flavilabris may have colonized the southeast of Central Europe. A migration of Adriatomediterranean faunal elements from the northwest Balkan Peninsula to Central Europe might be of more significance than previously assumed.

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Research Article Mon, 30 Aug 2021 16:47:08 +0300
Revision of Iranian Schoenlandella Cameron, 1905 (Hymenoptera, Braconidae, Cardiochilinae) with descriptions of two new species from Hormozgan province https://dez.pensoft.net/article/69090/ Deutsche Entomologische Zeitschrift 68(2): 261-268

DOI: 10.3897/dez.68.69090

Authors: Ilgoo Kang, Ali Ameri, Michael J. Sharkey

Abstract: The Iranian species of the genus Schoenlandella Cameron, 1905 are revised based on morphological characters. The current work provides a key to species of Iranian Schoenlandella and species descriptions of two new species: S. angustigena Kang sp. nov. and S. latigena Kang sp. nov. This work results in increasing the number of Iranian Schoenlandella species from one to three species.

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Research Article Mon, 30 Aug 2021 12:43:58 +0300
The Pyrenean species of Chelidura (Dermaptera, Forficulidae) https://dez.pensoft.net/article/68020/ Deutsche Entomologische Zeitschrift 68(2): 235-248

DOI: 10.3897/dez.68.68020

Authors: Pilar Jurado-Angulo, Yolanda Jiménez-Ruiz, Mario García-París

Abstract: The Pyrenees are inhabited by scattered populations of earwigs of the genus Chelidura Latreille, 1825. There is some controversy about the specific assignment of these populations: while most authors assign them to C. pyrenaica (Gené, 1832), other consider that C. aptera (Mégerlé, 1825) is also present in the Pyrenees. The main objective of this work was to revise the identity and synonyms of Pyrenean Chelidura. Specimens from recent fieldwork and collections (MNCN-CSIC) were used for morphological and molecular studies (cytochrome oxidase 1). All Pyrenean specimens shared similar cox1 sequences, very divergent from those of Alpine C. aptera. As a consequence, the variability observed in male cerci morphology from the Pyrenees, ranging from long and slightly curved to short and very curved, corresponded to C. pyrenaica, and the presence of C. aptera in the Pyrenees can be rejected. As previously suggested by Maccagno (1933) and Fontana et al. (2021), the revision of the synonymic list uncovered the misplacement of the name F. simplex Germar, 1825 under the synonymy of C. aptera, while it rather represents a synonym of C. pyrenaica (syn. nov.). Forficula simplex has nomenclatural priority over C. pyrenaica, however both names meet the requirements of the article 23.9.1 of the International Code of Zoological Nomenclature to retain the prevailing usage of C. pyrenaica (nomen protectum) over F. simplex (nomen oblitum). Additionally, we discuss the taxonomic status of Chelidura arverna David & Van Herrewege, 1973 stat. nov. from the French Massif Central.

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Research Article Tue, 10 Aug 2021 09:53:22 +0300
A third species of the rare ant genus Rotastruma (Hymenoptera, Formicidae) from Cambodia, with illustrated generic diagnoses and key to species https://dez.pensoft.net/article/63478/ Deutsche Entomologische Zeitschrift 68(2): 225-233

DOI: 10.3897/dez.68.63478

Authors: Shingo Hosoishi, Seiki Yamane, Heng Sokh

Abstract: The myrmicine genus Rotastruma is a rarely collected arboreal ant group and only two species, R. recava Bolton and R. stenoceps Bolton, are known from the Oriental Region. A new species, R. epispina sp. nov., is described from Cambodia based on the worker and queen castes. The new species is distinguished from R. recava and R. stenoceps by having the propodeal spine directed upwards and elongated peduncle of the petiole. Herein, we provide an update to the identification key and diagnostic characters for the genus Rotastruma.

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Research Article Mon, 5 Jul 2021 08:45:54 +0300
Balticeler kerneggeri gen. nov., sp. nov., an enigmatic Baltic amber fossil of the ground beetle subfamily Trechinae (Coleoptera, Carabidae) https://dez.pensoft.net/article/66181/ Deutsche Entomologische Zeitschrift 68(1): 207-224

DOI: 10.3897/dez.68.66181

Authors: Joachim Schmidt, Stephan Scholz, David R. Maddison

Abstract: Balticeler kerneggeri gen. nov., sp. nov., is described based on six fossil specimens preserved in Eocene Baltic amber and imaged using light microscopy and X-ray micro-computed tomography. Based on certain characters observed in the fossil species it is considered a “middle grade” Carabidae, outside of the large family Harpalinae (as it possesses a scrobal seta, the lack of which is a synapomorphy of that subfamily), but possessing four synapomorphies that indicate Balticeler belongs to a large clade of carabids including Harpalinae (anisochaetous Grade B antennal cleaner, conjunct mesocoxae, closed procoxal cavities, and a well-developed external lobe of the metepimeron). This remarkable beetle has several striking features, including lack of externally-visible sexually dimorphic characters, lack of lateral borders on the pronotum, and very long and thin mandibles and maxillae. In combination, these states are unique within Carabidae. We consider the presence of a dorsally completely open aedeagal median lobe as a synapomorphy of the fossil species with the subfamily Trechinae, a pubescent and relatively long second antennomere and a 4+2+2 pattern of umbilicate setae as synapomorphies of the supertribe Trechitae, and a quadrisetose clypeus as a synapomorphy with the Trechitae clade Bembidarenini + Trechini sensu Maddison et al. (2019). As it lacks a synapomorphy of Bembidarenini + Trechini, we propose that it is a member of the stem group of that clade.

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Research Article Wed, 2 Jun 2021 08:55:53 +0300
Four new species of Agraphydrus Régimbart, 1903 with additional faunistic record from China (Coleoptera, Hydrophilidae, Acidocerinae) https://dez.pensoft.net/article/66200/ Deutsche Entomologische Zeitschrift 68(1): 189-205

DOI: 10.3897/dez.68.66200

Authors: Zhen-ming Yang, Fenglong Jia, Lu Jiang, Qiang Guo

Abstract: Four new species of Agraphydrus Régimbart are described from China: A. pseudoniger sp. nov. from Shangyou County, Jiangxi Province, A. komareki sp. nov. from Shangchuan island, Taishan County, Guangdong Province, A. sabulosus sp. nov. from Fengkai, Guangdong Province, A. dapengensis sp. nov. from Dapeng peninsula, Shenzhen, Guangdong Province. Diagnosis and illustration of the new species are provided. The key given by Komarek and Hebauer (2018) to Chinese species of Agraphydrus Régimbart is updated.

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Research Article Tue, 18 May 2021 15:32:18 +0300
First instar nymphs of two peltoperlid stoneflies (Insecta, Plecoptera, Peltoperlidae) https://dez.pensoft.net/article/65540/ Deutsche Entomologische Zeitschrift 68(1): 179-188

DOI: 10.3897/dez.68.65540

Authors: Shodo Mtow, Tadaaki Tsutsumi

Abstract: The first instar nymphs of two peltoperlid stoneflies, i.e., Microperla brevicauda Kawai, 1958 of Microperlinae and Yoraperla uenoi (Kohno, 1946) of Peltoperlinae, were examined and described. Additionally, the phylogeny and groundplan of the first instar nymphs of Peltoperlidae and Plecoptera were considered. The first instar nymphs of M. brevicauda have a slender body with a prognathous head of typical shape; they represent a groundplan in Plecoptera. On the other hand, the first instar nymphs of Y. uenoi have a broad, cockroach-like body with an orthognathous and shortened head, the latter being regarded as a potential autapomorphy of Peltoperlinae. Such differences in body shape between the subfamilies are speculated to arise from heterochrony. The three-segmented cerci of Y. uenoi are characteristic to Systellognatha, whereas the four-segmented cerci of M. brevicauda were independently acquired within Microperlinae. The structure and distribution pattern of chloride cells in the first instar nymphs of Plecoptera were also discussed. The presence of coniform chloride cells is a potential groundplan of Arctoperlaria. One to two pairs of chloride cells are distributed on the first nine abdominal segments of M. brevicauda; this represents a groundplan character of Systellognatha. On the other hand, one to four pairs of chloride cells are found on the second to ninth abdominal segments of Y. uenoi; this distribution pattern may be an apomorphic groundplan of Peltoperlinae.

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Research Article Fri, 7 May 2021 16:55:56 +0300
Kazukuru gen. nov. – a new Ricaniidae planthopper from Solomon Islands (Hemiptera, Fulgoromorpha) https://dez.pensoft.net/article/63635/ Deutsche Entomologische Zeitschrift 68(1): 165-177

DOI: 10.3897/dez.68.63635

Authors: Adam Stroiński

Abstract: A new monotypic genus of ricaniid planthoppers (Hemiptera: Fulgoromorpha: Ricaniidae) from New Georgia Island (Solomon Islands), Kazukuru gen. nov., is described for K. zingiberis sp. nov. (type species). Habitus, female, external and internal genital structures of the new species are described and illustrated.

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Research Article Wed, 31 Mar 2021 15:58:03 +0300
Morphology, pollen preferences and DNA-barcoding of five Austrian species in the Colletes succinctus group (Hymenoptera, Apidae) https://dez.pensoft.net/article/55732/ Deutsche Entomologische Zeitschrift 68(1): 101-138

DOI: 10.3897/dez.68.55732

Authors: Katharina Zenz, Herbert Zettel, Michael Kuhlmann, Harald W. Krenn

Abstract: Most species of the Colletes succinctus group sensu Noskiewicz, 1936 are taxonomically uncertain. This study has chosen an integrative approach, including pollen analysis, morphology, male genitalia, morphometry, cuticle sculpture and DNA-barcoding (CO1) to investigate the five species that were reported from Austria. It includes a detailed analysis of the male genitalia and the first description of the C. pannonicus male. A syntype male from the island of Crete was designated as the lectotype of Colletes succinctus brevigena Noskiewicz, 1936 to fix the species identity. New distinguishing characters were found: in females the shape of the dorsal end of the fovea facialis and, in both sexes, the structure of maxillary palpi, as well as the different puncturation on the mesopleura. Unknown structures on sterna and genitalia of the males proved to be reliable morphological characters. An identification key is provided for all studied species. Morphometry of females did not allow a clear distinction of species. CO1 sequencing confirmed previous studies that only C. collaris clearly deviates from the other species, including C. pannonicus that was analysed for the first time. Pollen analysis showed polylectic, as well as oligolectic, pollen-collecting behaviour. The collected pollen of C. pannonicus confirmed the field observations that this species is strictly oligolectic on Tripolium pannonicum. Due to pronounced intraspecific variation, it is assumed that the species of the C. succinctus group are either species in statu nascendi or very young species. Therefore, it remains important to include ecological data in species identification.

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Research Article Tue, 2 Feb 2021 16:14:43 +0200
Biogeographical and evolutionary aspects of a Guineo-Congolian bushcricket tribe: Revision of the genera Cestromoecha Karsch, 1893 and Poreuomena Brunner von Wattenwyl, 1878, with the description of new species (Orthoptera, Tettigoniidae, Phaneropterinae) https://dez.pensoft.net/article/60193/ Deutsche Entomologische Zeitschrift 68(1): 45-79

DOI: 10.3897/dez.68.60193

Authors: Claudia Hemp, Bruno Massa

Abstract: The genera Cestromoecha and Poreuomena of the tribe Poreuomenini in Phaneropterinae are revised and new generic characters are given for both genera, and six new species are described in Poreuomena. The newly described species are P. biaculeata sp. nov., P. eala sp. nov., P. gracilicercata sp. nov., P. ivoriana sp. nov., P. matthaei sp. nov., and P. tshuapa sp. nov. Based on characters defining the two genera, three species so far listed under Cestromoecha are transferred to Poreuomena: P. crassipes Karsch, 1890, P. laeglae (Massa, 2015), and P. magnicerca (Massa, 2013). One species of Cestromoecha, C. mundamensis Karsch, 1896, is synonymised with C. tenuipes (Karsch, 1890) since no morphological differences were detected between the type specimens. Thus, two species remain with Cestromoecha, and Poreuomena now contains 16 species. Morphological closely-related species of Poreuomena suggest rapid speciation in the Congo Basin due to several expansions and shrinkages of the Guineo-Congolian forest belt since the Oligocene. At least two different morphological lineages are discernible. On the other hand the genus Cestromoecha Karsch, 1893 is a species-poor taxon.

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Research Article Tue, 5 Jan 2021 16:49:00 +0200
Review of the world species of Paroplitis Mason, 1981 (Hymenoptera, Braconidae, Microgastrinae), with description of three new species https://dez.pensoft.net/article/59641/ Deutsche Entomologische Zeitschrift 68(1): 33-43

DOI: 10.3897/dez.68.59641

Authors: Shunpei Fujie, George Japoshvili, Jose Fernandez-Triana

Abstract: The world species of the microgastrine genus Paroplitis (Hymenoptera: Braconidae) are revised. Three new species are described, P. horticola Fujie & Fernandez-Triana, sp. nov. and P. japonicus Fujie & Fernandez-Triana, sp. nov. from Japan and P. kakhetiensis Fujie, Japoshvili & Fernandez-Triana, sp. nov. from Georgia. P. vietnamensis van Achterberg & Fernandez-Triana, 2013 is re-described, based on additional specimens. P. wesmaeli Ruthe, 1860 is recorded from Georgia for the first time. A key to the nine known species (eight described and one undescribed) of the genus is provided.

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Research Article Tue, 5 Jan 2021 14:34:54 +0200
Comparative morphology of the musculature of the sting apparatus in Ampulex compressa (Hymenoptera, Ampulicidae) and Sceliphron destillatorium (Hymenoptera, Sphecidae) https://dez.pensoft.net/article/58217/ Deutsche Entomologische Zeitschrift 68(1): 21-32

DOI: 10.3897/dez.68.58217

Authors: Stefan Graf, Maraike Willsch, Michael Ohl

Abstract: The sting apparatus of aculeate Hymenoptera is derived from the ovipositor and is their most prominent apomorphy. In contrast to the frequently analysed sclerites of the sting apparatus, the associated musculature has largely been neglected. In this study, we use micro-computed tomography to present a detailed description of the musculature of the sting apparatus of Ampulex compressa (Ampulicidae) and Sceliphron destillatorium (Sphecidae). We found that 12 of 15 muscles corresponding to the sting apparatus are homologous between both species examined and 13 muscles in comparison with Hymenoptera described in the literature. All muscles identified as critical for the act of stinging were found in both species. Moreover, we found the ventral tergum 8-tergum 9 muscle and the tergum 8-tergum 8 muscles in A. compressa and the second valvifer-second valvifer muscle in S. destillatorium. For the first time, we describe the ventral tergum 8-tergum 9 muscle and the second valvifer-second valvifer muscle that interconnects both body sides, in Hymenoptera.

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Research Article Tue, 5 Jan 2021 13:41:00 +0200
Philibaetis gen. nov., a new genus from the Philippines (Ephemeroptera, Baetidae) https://dez.pensoft.net/article/59462/ Deutsche Entomologische Zeitschrift 68(1): 1-20

DOI: 10.3897/dez.68.59462

Authors: Thomas Kaltenbach, Jhoana M. Garces, Jean-Luc Gattolliat

Abstract: Investigations of type material and new material from the Philippines (Luzon) revealed that Baetis luzonensis Müller-Liebenau, 1982 and B. realonae Müller-Liebenau, 1982 do not belong to Baetis Leach, 1815. A new genus, Philibaetis gen. nov., is described to accommodate both species and both are re-described based on larvae. The new genus is characterised by having a rectangular labrum with a submarginal row of long, simple setae on the dorsal surface and ventrally on lateral margins long, simple, spine-like setae, on anterolateral margins long, feathered setae and medially long, bifid setae and a partial, submarginal row of lanceolate setae. Both mandibles have blade-like incisors and dorsally, a mediolateral patch of long, spine-like setae; additionally, the left mandible has a tuft of long, partly branched setae at the base of the subtriangular process. Philibaetis gen. nov. is further characterised by a hypopharynx with a medial tuft of stout setae and anterolaterally, two smaller tufts of stout setae, a galea-lacinia with the distal denti-seta tooth-like and directed against canines, a fore femur apically with stout setae, both on anterior and posterior side and without a femoral patch and a claw with one row of denticles and two or three subapical setae. The protogonostyli under the cuticle of male last instar larvae are folded in the Labiobaetis type, excluding their affiliation to the genus Baetis. COI sequences were obtained from both species. The genetic distance (Kimura 2-parameter) between them is 17.5% on average. Very limited genetic distances of 0% to 3% (0.75% on average) were found between specimens of P. luzonensis comb. nov.

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Research Article Tue, 5 Jan 2021 12:19:29 +0200
The subgenera Glabrobracon Fahringer, Lucobracon Fahringer and Uncobracon Papp of the genus Bracon Fabricius (Hymenoptera, Braconidae, Braconinae) in China, with the description of eleven new species https://dez.pensoft.net/article/57668/ Deutsche Entomologische Zeitschrift 67(2): 209-252

DOI: 10.3897/dez.67.57668

Authors: Yang Li, Jun-hua He, Xue-xin Chen

Abstract: The species of three subgenera of the genus Bracon Fabricius (Hymenoptera, Braconidae, Braconinae), i.e. Glabrobracon Fahringer, 1927; Lucobracon Fahringer, 1927; and Uncobracon Papp, 1996 from China are revised and 31 species are recognised, including 11 new species, i.e. Bracon (Glabrobracon) indistinctus sp. nov., B. (G.) leptotes sp. nov., B. (G.) longistriatus sp. nov., B. (G.) megaventris sp. nov., B. (Lucobracon) brevicarinatus sp. nov., B. (L.) coarctatus sp. nov., B. (L.) curculiovorus sp. nov., B. (L.) flavitestaceus sp. nov., B. (L.) quadratus sp. nov., B. (Uncobracon) eurysulcatus sp. nov. and B. (U.) longwangshanensis sp. nov. The new species are described and illustrated. Keys to the subgenera of the genus Bracon and the Chinese species of three subgenera Glabrobracon, Lucobracon and Uncobracon are provided.

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Research Article Thu, 19 Nov 2020 08:38:05 +0200
A DNA barcode library for ground beetles of Germany: the genus Agonum Bonelli, 1810 (Insecta, Coleoptera, Carabidae) https://dez.pensoft.net/article/56163/ Deutsche Entomologische Zeitschrift 67(2): 197-207

DOI: 10.3897/dez.67.56163

Authors: Michael J. Raupach, Karsten Hannig, Jérome Morinière, Lars Hendrich

Abstract: The ground beetle genus Agonum Bonelli, 1810 is a large genus of the tribe Platynini with many species that show high amounts of intraspecific variations, making a correct identification challenging. As part of the German Barcode of Life initiative, this publication provides a comprehensive DNA barcode library for species of Agonum that are reported for Germany. In total, DNA barcodes from 258 beetles and 23 species were analysed using the Barcode of Life Data System (BOLD) workbench, including sequences from former studies and 68 newly-generated sequences. The neighbour-joining analyses, based on K2P distances, revealed distinct clustering for all studied species, with unique Barcode Index Numbers (BINs) for 15 species (65%). BIN sharing but distinct clustering was found for three species pairs: Agonum micans/Agonum scitulum, Agonum impressum/Agonum sexpunctatum and Agonum duftschmidi/Agonum emarginatum. The given dataset and its analysis represent another important step in generating a comprehensive DNA barcode library for the ground beetles of Germany and Central Europe in terms of modern biodiversity research.

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Research Article Mon, 19 Oct 2020 17:00:53 +0300
Revision of the Plagiolepis schmitzii group with description of Pl. invadens sp. nov. – a new invasive supercolonial species (Hymenoptera: Formicidae) https://dez.pensoft.net/article/53199/ Deutsche Entomologische Zeitschrift 67(2): 183-196

DOI: 10.3897/dez.67.53199

Authors: Bernhard Seifert

Abstract: Using high-resolution stereomicroscopy and exploratory data analyses, a taxonomic revision of the cryptic species close to Plagiolepis schmitzii Forel, 1895, called Pl. schmitzii group, was conducted. Morphology was numerically recorded under highly standardised conditions considering absolute size and 16 shape, pubescence and surface characters. A key to the non-parasitic Westpalaearctic species of the ant genus Plagiolepis Mayr, 1861 is provided which firstly separates, on species group level, the Pl. pygmaea (Latreille) species group, the Pl. pallescens Forel species group and the Pl. schmitzii species group and, finally, on species level, the cryptic species of the latter group. The recognised species of the Pl. schmitzii species group are Pl. schmitzii Forel, 1895 (invasive species), Pl. barbara Santschi, 1911, Pl. atlantis Santschi, 1920 and Pl. invadens sp. nov. (invasive species) that is described as new from a supercolony in Germany. Based on morphological arguments, the taxa Pl. barbara var. madeirensis Emery, 1921, Pl. maura polygyna Santschi, 1922 and Pl. schmitzii var. tingitana Santschi, 1936 are recognised as junior synonyms of Pl. schmitzii, the taxa Pl. schmitzii crosi Santschi, 1920, Pl. pallescens var. kabyla Santschi, 1920 and Pl. perperamus Salata et al., 2018 as junior synonyms of Pl. atlantis and the taxon Pl. maura Santschi, 1920 as junior synonym of Pl. barbara. A concluding comparative section suggests that pre-adaptations for anthropogenous dispersal and transformation to supercoloniality in introduction areas are apparently common traits in Plagiolepis ants.

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Research Article Mon, 21 Sep 2020 15:16:35 +0300
Phylogeny and classification of the genus-group taxa of Loxandrina (Coleoptera, Carabidae, Abacetini) https://dez.pensoft.net/article/55985/ Deutsche Entomologische Zeitschrift 67(2): 151-182

DOI: 10.3897/dez.67.55985

Authors: Kipling Will

Abstract: Bayesian and parsimony phylogenetic analyses of combined and partitioned datasets of molecular (partial sequences of 28S, wg, COI, and CAD) and morphological (51 characters of adults) data for exemplar taxa of five outgroup and 76 ingroup abacetine carabids resulted in a monophyletic Loxandrina Erwin & Sims, 1984 that is split into Australian and American clades. The genus Loxandrus LeConte, 1853 as previously delimited is not monophyletic relative to numerous genus-level taxa in Abacetini Chaudoir, 1873 and is restricted to a subgenus of North American species. A reclassification and nomenclatural changes for the subtribe that are consistent with the phylogeny are provided. Three genera are removed from Loxandrina: Aulacopodus Britton, 1940 moved to Pterostichini Bonelli, 1810; Cosmodiscus Sloane, 1907 and Tiferonia Darlington, 1962 moved to Abacetina. Based on the phylogenetic relationships and nomenclatural priority only four genera are recognized in Loxandrina: Cerabilia Laporte, 1867, Zeodera Laporte, 1867, Pediomorphus Chaudoir, 1878, and Oxycrepis Reiche, 1843. All other previously recognized genera are treated as subgenera. The classification change created eight secondary homonyms that are resolved by the proposal of the following: Oxycrepis gebi, replacement name for O. balli (Straneo, 1993); O. amatona, replacement name for O. matoana (Straneo, 1993); O. xiproma, replacement name for O. proxima (Straneo, 1993); O. rasutulis, replacement name for O. suturalis (Straneo, 1993); O. laevinota, replacement name for O. laevicollis (Bates, 1871); O. arvulap, replacement name for O. parvula (Straneo, 1951); O. noaffine, replacement name for O. affinis (Straneo, 1991); O. alutona, replacement name for O. notula (Tschitschérine, 1901). An overview of the morphological characteristics and diagnostic features of Loxandrina taxa is provided. A key and habitus images are provided for identification of genera and subgenera. The possible historical biogeography of the group is discussed in light of their phylogenetic relationships and past geological events.

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Research Article Wed, 2 Sep 2020 14:23:40 +0300
Anchored between heaven and earth – a new flightless brown lacewing from Peru (Neuroptera, Hemerobiidae) https://dez.pensoft.net/article/56008/ Deutsche Entomologische Zeitschrift 67(2): 141-149

DOI: 10.3897/dez.67.56008

Authors: Ulrike Aspöck, Horst Aspöck, Axel Gruppe

Abstract: Male and female of Nusalala peruana sp. nov., a flightless hemerobiid from the Andes mountain range of northern Peru, at a height of almost 4000 m, are described, figured and documented as the first record of a brachypterous, flightless species of Nusalala Navás, 1913, from this country. The other two congeneric, brachypterous species are from high altitudes in Colombia and Costa Rica and have been described in the male sex only – the females remain unknown. The coriaceous domed forewings are shared by all three brachypterous Nusalala species. The ribbon-like hindwings of the male of N. peruana sp. nov. are unique, since those of the other brachypterous males are scale-like, as are the hindwings of the female of N. peruana sp. nov. Distribution and evolutionary backgrounds of brachyptery and flightlessness in Neuropterida are discussed.

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Research Article Fri, 14 Aug 2020 15:49:15 +0300
Recent additions to the list of German sawflies (Hymenoptera, Symphyta) https://dez.pensoft.net/article/54002/ Deutsche Entomologische Zeitschrift 67(2): 127-139

DOI: 10.3897/dez.67.54002

Authors: Andrew Liston, Marko Prous

Abstract: Two tenthredinid sawfly species are newly recorded in Germany: Pristiphora krausi (Lacourt, 2006) and P. melagonia sp. nov. The latter is also recorded from Greece and Spain. These species belong to the Pristiphora depressa group, as confirmed by genetic data. Additional data are presented for seven other tenthredinid species which have only rarely been recorded in Germany and for which previously-published data are incomplete. Identification keys to the West Palaearctic species of Eurhadinoceraea and the Pristiphora depressa group are included. Other species of Symphyta, first recorded in Germany since the publication of the most recent checklist, are listed in a table, with references to literature which describes identification characters.

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Research Article Tue, 21 Jul 2020 14:30:54 +0300
The Nepalese species of the genus Enicospilus Stephens, 1835 (Hymenoptera, Ichneumonidae, Ophioninae): a preliminary revision and identification key to species https://dez.pensoft.net/article/51332/ Deutsche Entomologische Zeitschrift 67(1): 69-126

DOI: 10.3897/dez.67.51332

Authors: So Shimizu

Abstract: A total of 10 species of Enicospilus (Hymenoptera, Ichneumonidae, Ophioninae) have previously been reported from Nepal. Six new species are described here (E. alleni Shimizu sp. nov., E. kakanicus Shimizu sp. nov., E. nepalensis Shimizu sp. nov., E. nikami Shimizu sp. nov., E. phulchokiensis Shimizu sp. nov., and E. tangi Shimizu sp. nov.), and 10 are newly recorded (E. ashbyi Ashmead, 1904, E. bifasciatus (Uchida, 1928), E. capensis (Thunberg, 1824), E. flavocephalus (Kirby, 1900), E. formosensis (Uchida, 1928), E. grammospilus (Enderlein, 1921), E. pudibundae (Uchida, 1928), E. purifenestratus (Enderlein, 1921), E. yonezawanus (Uchida, 1928), and E. zebrus Gauld & Mitchell, 1981) from Nepal. A preliminary identification key to the Nepalese species of Enicospilus is provided. The elevational pattern of Nepalese Enicospilus is briefly discussed. Enicospilus purifenestratus is also recorded for the first time from Brunei.

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Research Article Mon, 11 May 2020 09:27:28 +0300
A comparative description of the mesosomal musculature in Sphecidae and Ampulicidae (Hymenoptera, Apoidea) using 3D techniques https://dez.pensoft.net/article/49493/ Deutsche Entomologische Zeitschrift 67(1): 51-67

DOI: 10.3897/dez.67.49493

Authors: Maraike Willsch, Frank Friedrich, Daniel Baum, Ivo Jurisch, Michael Ohl

Abstract: Conflicting hypotheses about the relationships among the major lineages of aculeate Hymenoptera clearly show the necessity of detailed comparative morphological studies. Using micro-computed tomography and 3D reconstructions, the skeletal musculature of the meso- and metathorax and the first and second abdominal segment in Apoidea are described. Females of Sceliphron destillatorium, Sphex (Fernaldina) lucae (both Sphecidae), and Ampulex compressa (Ampulicidae) were examined. The morphological terminology provided by the Hymenoptera Anatomy Ontology is used. Up to 42 muscles were found. The three species differ in certain numerical and structural aspects. Ampulicidae differs significantly from Sphecidae in the metathorax and the anterior abdomen. The metapleural apodeme and paracoxal ridge are weakly developed in Ampulicidae, which affect some muscular structures. Furthermore, the muscles that insert on the coxae and trochanters are broader and longer in Ampulicidae. A conspicuous characteristic of Sphecidae is the absence of the metaphragma. Overall, we identified four hitherto unrecognized muscles. Our work suggests additional investigations on structures discussed in this paper.

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Research Article Mon, 11 May 2020 09:25:08 +0300
A review of Himalcercyon stat. nov., with description of a new species from the Chinese Himalaya and an updated key to Asian genera of Megasternini (Coleoptera, Hydrophilidae) https://dez.pensoft.net/article/50078/ Deutsche Entomologische Zeitschrift 67(1): 35-49

DOI: 10.3897/dez.67.50078

Authors: Fenglong Jia, Zulong Liang, Martin Fikáček

Abstract: Himalcercyon Hebauer, 2002 stat. nov. is elevated to genus rank based on the unique form of its mesoventral elevation. The genus is reviewed, redescribed, and illustrated in detail. Two species are recognized: Himalcercyon mirus (Hebauer, 2002) comb. nov. from Nepal and H. franzi sp. nov. from Chinese Himalaya (Xizang Autonomous Region). Both species are illustrated and diagnosed. An updated key to the Asian genera of the tribe Megasternini (Coleoptera, Hydrophilidae, Sphaeridiinae) is provided, along with the SEM micrographs of ventral morphology of these genera. New replacement name Oreosternum nom. nov. is proposed for Oreocyon Hebauer, 2002 which is preoccupied by Oreocyon Marsh, 1872 (Mammalia, Oxyenidae) and Oreocyon Krumbiegel, 1949 (Mammalia, Canidae).

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Research Article Mon, 11 May 2020 08:37:32 +0300
Molecular taxonomy of Tomares hairstreaks (Lepidoptera, Lycaenidae, Theclinae) https://dez.pensoft.net/article/50252/ Deutsche Entomologische Zeitschrift 67(1): 19-33

DOI: 10.3897/dez.67.50252

Authors: Vazrick Nazari, Wolfgang ten Hagen

Abstract: Tomares hairstreaks comprise about 10 species distributed from Europe and North Africa to Central Asia. The taxonomy of the genus is hampered by the absence of diagnostic characters by which specimens can be unambiguously assigned to species. Our investigation of morphology and DNA barcode variations within and between Tomares species shows that while well-defined species (T. ballus, T. mauritanicus, T. callimachus, T. desinens and T. fedtschenkoi) diverge, poorly characterized taxa (T. nogelii, T. nesimachus, T. dobrogensis, T. romanovi and T. telemachus) show very little to no differentiation in mtDNA. We reinstate Tomares callimachus spp. hafis (Kollar, 1849) as a valid subspecies (stat. rev.) and propose taxa telemachus Zhdanko, 2000 and uighurica Koçak, Seven & Kemal, 2000 as synonyms of T. romanovi and T. nogelii nogelii respectively (syn. nov.). We relegate Polyommatus epiphania Boisduval, 1848, recently revived as a valid subspecies of T. callimachus, back to synonymy under the latter, and reconsider the status of T. nogelii dobrogensis (Caradja, 1895) in the light of new molecular data. We use a nuclear gene (EF-1α) in addition to COI barcodes to reconstruct the phylogeny of the group.

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Research Article Tue, 5 May 2020 08:37:12 +0300
Biology of two European Tenthredo species (Hymenoptera, Tenthredinidae) feeding on Gentiana https://dez.pensoft.net/article/49741/ Deutsche Entomologische Zeitschrift 67(1): 13-18

DOI: 10.3897/dez.67.49741

Authors: Andrew Liston, Ewald Altenhofer, Romana Netzberger, Marko Prous

Abstract: Very few sawflies using Gentianaceae as larval host plants have been recorded. We identified larvae collected in Austria on Gentiana asclepiadea L. as Tenthredo atra Linnaeus, 1758 and T. propinqua Klug, 1817. If its current taxonomic circumscription as a single species is accepted, T. atra is a highly polyphagous species, whereas T. propinqua may be more specialised: Gentiana asclepiadea is its first recorded host. We sequenced plant DNA from the head of one T. propinqua larva, which confirmed that it had been feeding on this plant. This is the first recorded use of G. asclepiadea by sawfly larvae. Larvae are illustrated, and identification characters are described.

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Research Article Fri, 21 Feb 2020 10:41:07 +0200
The Tamarix feeding Leafhopper genus Opsius Fieber, 1866 (Hemiptera, Cicadellidae, Deltocephalinae, Opsiini) in the Kingdom of Saudi Arabia, with description of a new species https://dez.pensoft.net/article/46662/ Deutsche Entomologische Zeitschrift 67(1): 1-12

DOI: 10.3897/dez.67.46662

Authors: Saad A. El-Sonbati, Michael R. Wilson, Hathal M. Al Dhafer

Abstract: The leafhopper genus Opsius Fieber, 1866 is revised for the Kingdom of Saudi Arabia. Seven species are treated, including three that previously were reported by Dlabola (1979), O. pallasi (Lethierry, 1874), O. tigripes (Lethierry, 1876), and O. versicolor (Distant, 1908). Opsius heydeni (Lethierry, 1876), O. richteri Dlabola 1960, and O. scutellaris (Lethierry, 1874) are reported for the first time from the Kingdom. A new species, O. wilsoni El-Sonbati, sp. nov. is described from the southwestern region of the Kingdom of Saudi Arabia. A key to the species of Opsius of the Kingdom is also provided.

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Research Article Tue, 14 Jan 2020 09:32:39 +0200
Species composition and diagnoses of leaf- and fruit-scarring beetles (Coleoptera, Chrysomelidae) infesting bananas and plantains (Zingiberales, Musaceae) in the Indian subcontinent https://dez.pensoft.net/article/47447/ Deutsche Entomologische Zeitschrift 66(2): 179-202

DOI: 10.3897/dez.66.47447

Authors: Kaniyarikkal D. Prathapan, Janakiraman Poorani, S. Amritha Kumari, C. Anuradha, Balakrishnan Padmanaban, Ramalingam Thanigairaj

Abstract: Leaf- and fruit-feeding chrysomelids (Coleoptera) on bananas and plantains (Musaceae, Zingiberales) cause major losses to banana growers in the northern and northeastern regions of India, Bangladesh, and other parts of Southeast Asia. The species composition of these beetles has not been studied so far in India and wrong names or wrong name combinations in the literature have caused confusion. Most particularly, the Central and South American apecies of Colaspis hypochlora Lefèvre (Chrysomelidae, Eumolpinae) has been erroneously reported as occurring in India and Bangladesh, and this name has been used for the Indian species. Based on extensive surveys for leaf- and fruit-feeding chrysomelids in the northern and northeastern regions of India from 2015 to 2019, three species of banana-feeding chrysomelids, namely, Basilepta subcostata (Jacoby) (Eumolpinae), Bhamoina varipes (Jacoby), and a new species, Sphaeroderma cruenta sp. nov. (Galerucinae, Alticini), are documented. Of these, the latter two are recorded as pests of banana in India for the first time. An illustrated diagnostic account of these three species is given to facilitate their identification by economic entomologists. COI sequences of populations of B. subcostata from Assam and Uttar Pradesh showed 98–100% homology, indicating that these populations are conspecific and that COI sequences can be used for rapid species determination. Brief notes on the biology and available management options for these pests are also given.

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Research Article Fri, 13 Dec 2019 09:23:29 +0200
First known larva of omicrine genus Psalitrus d’Orchymont (Coleoptera, Hydrophilidae) https://dez.pensoft.net/article/34300/ Deutsche Entomologische Zeitschrift 66(1): 107-118

DOI: 10.3897/dez.66.34300

Authors: Yûsuke N. Minoshima

Abstract: The larval morphology of the water scavenger beetle Psalitrus yamatensis Hoshina & Satô, 2005 is described based on a specimen collected from Fukuoka Prefecture, Kyushu, Japan. This is the first description of the larval morphology of the hydrophilid genus Psalitrus d’Orchymont, 1919, as well as the first description of larval chaetotaxy of the tribe Omicrini. Species-level identification of the larva was performed using DNA barcoding of a molecular marker: a 658 bp fragment of the mitochondrial cytochrome oxidase I. A description including chaetotaxy of head capsule and head appendages, diagnosis, and illustrations of the larva is provided. Psalitrus larvae can be distinguished from other known larvae of the tribe Omicrini by the morphology of the head and legs. The larva shares characters with other known larvae of Omicrini; potential plesiomorphies are shared with Cylominae and aquatic hydrophilids; some characters are also shared with larvae of Megasternini and Sphaeridiini.

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Research Article Tue, 25 Jun 2019 09:15:36 +0300
Contribution to the knowledge of male and female eremochaetid flies in the late Cretaceous amber of Burma (Diptera, Brachycera, Eremochaetidae) https://dez.pensoft.net/article/33914/ Deutsche Entomologische Zeitschrift 66(1): 75-83

DOI: 10.3897/dez.66.33914

Authors: Qingqing Zhang, Junfeng Zhang

Abstract: A new and a previously known species of the genus Zhenia Q. Zhang, 2016 (Eremochaetidae) are illustrated and described based on two males and a female in amber: Zhenia burmensis sp. nov. and Z. xiai Q. Zhang, 2016. The male Z. xiai is the first male of this species recorded. The relationships of Archisargoidea (including Eremochaetidae, Zhenia) are reassessed based on male genitalia. The superfamily is more likely related to the Stratiomyomorpha than to the Muscomorpha (including Nemestrinoidea). The components and structures of the ovipositor are re-illustrated. The results of our comparative study demonstrate that the ovipositor of Zhenia is similar in shape and detail to that of Rhagoletis pomonella (Walsh, 1867) (Tephritidae). This study concludes that the ovipositor of Zhenia is most likely formed from abdominal eighth and ninth segments instead of the cerci, as a previous study found.

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Research Article Fri, 14 Jun 2019 10:05:35 +0300
On Bulgarian sawflies, including a new species of Empria (Hymenoptera, Symphyta) https://dez.pensoft.net/article/34309/ Deutsche Entomologische Zeitschrift 66(1): 85-105

DOI: 10.3897/dez.66.34309

Authors: Andrew Liston, Marko Prous, Jan Macek

Abstract: Thirty-nine species of sawfly (Symphyta) are recorded for the first time in Bulgaria. Most of these were collected during early spring of 2018, in the south-east of the country (Burgas and Varna Provinces). Empria aridicola Macek & Prous, sp. nov. is described as new to science from specimens collected in several central, east and south European countries. Lectotypes are designated for Poecilosoma parvula Konow, 1892, Empria pravei Dovnar-Zapolskij, 1925 and E. pseudoklugi Dovnar-Zapolskij, 1929. Empria pravei and Sciapteryx byzantina Benson, 1968 are at present only known in Europe from the coastal zone of the Black Sea. The new Bulgarian records of Hoplocampa cantoti Chevin, 1986 and Neomessa steusloffi (Konow, 1892) represent large extensions in their recorded ranges, previously comprising respectively only northern France, and north-eastern Germany. Possible host plant associations are noted for several species, based on observations of adults.

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Research Article Fri, 14 Jun 2019 09:24:26 +0300
The Rubens morph of Formica exsecta Nylander, 1846 and its separation from Formica fennica Seifert, 2000 (Hymenoptera, Formicidae) https://dez.pensoft.net/article/34868/ Deutsche Entomologische Zeitschrift 66(1): 55-61

DOI: 10.3897/dez.66.34868

Authors: Bernhard Seifert

Abstract: A study of numeric morphology-based alpha-taxonomy (NUMOBAT) considering the species Formica exsecta Nylander, 1846 and F. fennica Seifert, 2000 was performed in 166 nest samples with 485 worker individuals originating from 117 localities of the Palaearctic west of 59°E. The presence of intraspecific pilosity dimorphism is shown for F. exsecta. The setae-reduced phenotype, termed the Rubens morph, shows a frequency of about 25%, and the more abundant setae-rich phenotype, termed the Normal morph, one of 75%. The frequency of nests containing workers of both phenotypes is 15.5% in 58 samples from Denmark, Sweden, and Finland. Applying the DIMORPH test of Seifert (2016) on this territory, it is demonstrated that the association of Rubens and Normal phenotypes within the same nest cannot be interpreted as parabiosis of independent species (p=0.017) or as temporary (p=0.0004) and permanent (p=0.0001) socially parasitic association, whereas genetically mediated intraspecific dimorphism is most likely (p=0.659, all p data according to Fisher’s exact test). The Rubens morph of F. exsecta is phenotypically most similar to F. fennica but is safely separable by four different forms of exploratory data analyses using nest centroids (NC) as input data: NC-Ward, NC-part.hclust, NC-part.kmeans, and NC-NMDS-k-means. Data on zoogeography and the narrow climate niche indicate that F. fennica is unlikely to occur in Norway.

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Research Article Wed, 29 May 2019 16:11:23 +0300
About Notiophilus Duméril, 1806 (Coleoptera, Carabidae): Species delineation and phylogeny using DNA barcodes https://dez.pensoft.net/article/34711/ Deutsche Entomologische Zeitschrift 66(1): 63-73

DOI: 10.3897/dez.66.34711

Authors: Michael J. Raupach, Karsten Hannig, Jérome Morinière, Lars Hendrich

Abstract: The genus Notiophilus Duméril, 1806 is a distinctive taxon of small, diurnal and morphologically similar beetles exhibiting large eyes and widened second elytral intervals. In this study we analysed the effectiveness of DNA barcodes to discriminate 67 specimens that represent 8 species of Notiophilus from Central Europe. Interspecific K2P distances below 2.2% were found for N. biguttatus (Fabricius, 1779) and N. quadripunctatus Dejean, 1826, whereas intraspecific distances with values > 2.2% were revealed for N. rufipes Curtis, 1829. An additional phylogenetic analysis of all available species revealed a close relationship of N. directus Casey, 1920, N. semistriatus Say, 1823, N. simulator Fall, 1906 and N. sylvaticus Dejean, 1831, possibly indicating a radiation of these species in North America. Low support values of most other nodes, however, do not allow additional phylogenetic conclusions.

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Research Article Wed, 29 May 2019 15:25:55 +0300
Type catalogue of the thick-headed flies (Diptera, Conopidae) in the collection of the Museum für Naturkunde Berlin https://dez.pensoft.net/article/33814/ Deutsche Entomologische Zeitschrift 66(1): 41-53

DOI: 10.3897/dez.66.33814

Authors: Jens-Hermann Stuke, Joachim Ziegler

Abstract: Type material of thick-headed flies (Diptera, Conopidae) in the collection of the Museum für Naturkunde Berlin (ZMHB) is documented. The entire collection holds primary type material (i.e. holotypes, lectotypes, syntypes) of 73 species. Five Conops species with previously unknown subgenus belong to the subgenus Asiconops: C. frontosus Kröber, 1916; C. indicus Kröber, 1916; C. maculiventris Kröber, 1916; C. nigrofasciatus Kröber, 1916; and C. punctifrons Kröber 1916. Two new synonyms are introduced: Conops vaginalis Rondani, 1865 syn. nov. of Conops truncatus Loew, 1847 and Siniconops fuscatus Qiao & Chao, 1998 syn. nov. of Physocephala sepulchralis Brunetti, 1912.

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Research Article Wed, 15 May 2019 12:19:33 +0300
Redescriptions of thirteen species of chewing lice in the Brueelia-complex (Phthiraptera, Ischnocera, Philopteridae), with one new synonymy and a neotype designation for Nirmus lais Giebel, 1874 https://dez.pensoft.net/article/32423/ Deutsche Entomologische Zeitschrift 66(1): 17-39

DOI: 10.3897/dez.66.32423

Authors: Daniel R. Gustafsson, Lucie Oslejskova, Tomas Najer, Oldrich Sychra, Fasheng Zou

Abstract: Thirteen species of chewing lice in the Brueelia-complex are redescribed and illustrated. They are: Brueelia blagovescenskyi Balát, 1955, ex Emberiza schoeniclus (Linnaeus, 1758); B. breueri Balát, 1955, ex Chloris chloris (Linnaeus, 1758); B. conocephala (Blagoveshchensky, 1940) ex Sitta europaea (Linnaeus, 1758); B. ferianci Balát, 1955, ex Anthus trivialis (Linnaeus, 1758); B. glizi Balát, 1955, ex Fringilla montifringilla Linnaeus, 1758; B. kluzi Balát, 1955, ex Fringilla coelebs Linnaeus, 1758; B. kratochvili Balát, 1958, ex Motacilla flava Linnaeus, 1758; B. matvejevi Balát, 1981, ex Turdus viscivorus Linnaeus, 1758; B. pelikani Balát, 1958, ex Emberiza melanocephala Scopoli, 1769; B. rosickyi Balát, 1955, ex Sylvia nisoria (Bechstein, 1792); B. vaneki Balát, 1981, ex Acrocephalus schoenobaenus (Linnaeus, 1758); Guimaraesiella haftorni (Balát, 1958) ex Turdus iliacus Linnaeus, 1758; G. lais (Giebel, 1874) ex Luscinia megarhynchos (Brehm, 1831). Redescriptions are made from type material where available. Holotypes are identified in Balát’s material when possible, and lectotypes are designated for B. blagovescenskyi, B. breueri, B. glizi, B. ferianci, B. kluzi, B. kratochvili, B. pelikani, and B. rosickyi; a neotype of Nirmus lais Giebel, 1874 is designated. Brueelia weberi Balát, 1982, is placed as a synonym of Brueelia conocephala (Blagoveshchensky, 1940).

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Research Article Mon, 4 Feb 2019 09:58:30 +0200
Playing hard to get: two new species of subterranean Trechini beetles (Coleoptera, Carabidae, Trechinae) from the Dinaric Karst https://dez.pensoft.net/article/31754/ Deutsche Entomologische Zeitschrift 66(1): 1-15

DOI: 10.3897/dez.66.31754

Authors: Roman Lohaj, Teo Delić

Abstract: Almost 200 years of continuous and systematic research in subterranean habitats of the Dinaric Karst and adjoining areas have resulted in the discovery of more than 400 specialized subterranean beetles. Among these, a special place belongs to the morphologically well distinguished and elusive, so called aphaenopsoid trechine beetles, which are characterized by a prolonged head, pronotum and appendages, and widened, ovoid-shaped elytra. Two new species of aphaenopsoid trechines – Derossiella lukici sp. n. from two deep pits on Mt Biokovo, Croatia, and Adriaphaenops petrimaris sp. n. from Pištet 4 Cave, Kameno more, Montenegro – are described, illustrated, and compared with closely related congeners. Identification keys for both genera and an annotated catalogue for all Adriaphaenops species, as well as data on the distribution and the ecology of these remarkable species, are provided and discussed.

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Research Article Mon, 21 Jan 2019 09:39:18 +0200
Revision of Zosteragathis Sharkey of Thailand (Hymenoptera, Braconidae, Agathidinae, Agathidini) https://dez.pensoft.net/article/25772/ Deutsche Entomologische Zeitschrift 65(2): 225-253

DOI: 10.3897/dez.65.25772

Authors: Michael J. Sharkey, Eric G. Chapman

Abstract: Based on cladistic analyses recently conducted by Sharkey and Chapman, the genus Zosteragathis Sharkey is revised. Twenty-two species are reported from Thailand, three previously described species, Z. samensis, Z. contrasta and Z. nuichuaensis, and 19 new species, i.e., Z. chaiyaphumensis, Z. eukos, Z. hinensis, Z. hongensis, Z. inthanonensis, Z. krachanensis, Z. lampangensis, Z. lampooensis, Z. luangensis, Z. ngamensis, Z. perknos, Z. petchaburiensis, Z. phahompokensis, Z. phuphanensis, Z. sakaeratensis, Z. sakonensis, Z. samensis, Z. surinensis, Z. taemensis, Z. tonensis. Members of Zosteragathis are known from the Australian, Ethiopian, Oceania, Oriental, and eastern Palaearctic regions.

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Research Article Tue, 14 Aug 2018 15:18:09 +0300
Revision of the Quedius fauna of Middle Asia (Coleoptera, Staphylinidae, Staphylininae) https://dez.pensoft.net/article/27033/ Deutsche Entomologische Zeitschrift 65(2): 117-159

DOI: 10.3897/dez.65.27033

Authors: Maria Salnitska, Alexey Solodovnikov

Abstract: Twenty eight species of the genus Quedius from Middle Asia comprising Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan and Uzbekistan, are revised. Quedius altaicus Korge, 1962, Q. capitalis Eppelsheim, 1892, Q. fusicornis Luze, 1904, Q. solskyi Luze, 1904 and Q. cohaesus Eppelsheim, 1888 are redescribed. The following new synonymies are established: Q. solskyi Luze, 1904 = Q. asiaticus Bernhauer, 1918, syn. n.; Q. cohaesus Eppelsheim, 1888 = Q. turkmenicus Coiffait, 1969, syn. n., = Q. afghanicus Coiffait, 1977, syn. n.; Q. hauseri Bernhauer, 1918 = Q. peneckei Bernhauer, 1918, syn. n., = Q. ouzbekiscus Coiffait, 1969, syn. n.; Q. imitator Luze, 1904 = Q. tschinganensis Coiffait, 1969, syn. n.; Q. novus Eppelsheim, 1892 = Q. dzambulensis Coiffait, 1967, syn. n., Q. pseudonigriceps Reitter, 1909 = Q. kirklarensis Korge, 1971, syn. n. Lectotypes are designated for Q. asiaticus Bernhauer, 1918, Q. fusicornis Luze, 1904, Q. hauseri Bernhauer, 1918, Q. imitator Luze, 1904, Q. novus Eppelsheim, 1892 and Q. solskyi Luze, 1904. For all revised species, taxonomy, distribution and bionomics are summarized. Quedius fuliginosus (Gravenhorst, 1802), Q. sundukovi Smetana, 2003 and Q. pseudonigriceps Reitter, 1909 are recorded for Middle Asia for the first time. One species from the Q. coloratus-group, found to be new to science is not described due to shortage of material. Another possibly new species is tentatively identified as Q. fulvicollis Stephens, 1833 until the taxonomy of that widespread species is revised. An identification key to all species is provided.

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Research Article Tue, 31 Jul 2018 10:05:58 +0300
Review of the flower-inhabiting water scavenger beetle genus Cycreon (Coleoptera, Hydrophilidae), with descriptions of new species and comments on its biology https://dez.pensoft.net/article/26261/ Deutsche Entomologische Zeitschrift 65(1): 99-115

DOI: 10.3897/dez.65.26261

Authors: Emmanuel Arriaga-Varela, Sin Yeng Wong, Alexander Kirejtshuk, Martin Fikácek

Abstract: The hydrophilid genus Cycreon Orchymont, 1919, previously known from two historical specimens only, is reviewed based on the numerous material collected recently from the inflorescences of various Araceae species in the Malay Peninsula and Borneo. Four species are recognized in the genus: C. sculpturatus Orchymont, 1919 from Sumatra, C. armandi Shatrovskiy, 2017 from Singapore, C. adolescens sp. n. from peninsular Malaysia, and C. floricola sp. n. with two subspecies, the nominotypical one from Peninsular Malaysia, and C. floricola borneanus subsp. n. from Borneo. All species are very similar, differing only by the pronotal punctation, shape of the clypeus and the mentum, and the form of the median lobe of the aedeagus. Specimens of C. floricola sp. n. and C. adolescens sp. n. were collected from inflorescences of various genera of the family Araceae. The field observations and analysis of mid gut contents indicates that they feed on organic material on internal organs of the inflorescences, including the pollen of the host plant. They were also observed to carry a large amount of pollen and are likely pollinators of their host species of Araceae.

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Research Article Tue, 12 Jun 2018 10:13:01 +0300
Anthropogenic dispersal of a snakefly (Insecta, Neuropterida) – a singular phenomenon or a model case in Raphidioptera? https://dez.pensoft.net/article/19859/ Deutsche Entomologische Zeitschrift 64(2): 123-131

DOI: 10.3897/dez.64.19859

Authors: Horst Aspöck, Ulrike Aspöck, Axel Gruppe, Marcia Sittenthaler, Elisabeth Haring

Abstract: The Mediterranean snakefly Raphidia mediterranea H. Aspöck, U. Aspöck & Rausch, 1977 – known from many parts of the Balkan Peninsula, several Aegean islands, southern parts of Italy, northwest of Anatolia and a few localities in Eastern Europe, yet not recorded in Central Europe – was surprisingly found with an astoundingly high population density on bushes in the yard of an old farmhouse at a comparatively high altitude (800 m) in Upper Austria, north of the Danube River, in 2013. This spectacular phenomenon was again observed in the following years (2014, 2015, 2016, 2017), and in 2016 the suspicion that the larvae develop in the straw of the thatched roof of the farmhouse could be confirmed by findings of larvae, pupae, and exuviae.It is most likely, that the occurrence of this Raphidia species in Austria is to be traced back to a human-caused introduction at some point in time. It remains, however, unknown when and specifically how this event might have occurred. Morphologically no substantial differences were found between specimens from Greece, Italy and Upper Austria. In addition, the genetic uniformity (using 3 genes: cox1, cox3, and 28S) of the populations was verified. This supports an earlier hypothesis that the occurrence of the species, as well as that of R. mediterranea in Italy, Anatolia and perhaps elsewhere, may be related to importation of goods involving wood or soil. A molecular genetic analysis of several Raphidia species confirmed the present morphology-based concept of their systematic position. The means of dispersal of Raphidioptera are largely unknown. We do not know of any other similar cases of anthropogenic dispersal of a snakefly, but it cannot be excluded that human activities may have played a greater role in the dispersal of Raphidioptera than previously assumed. Phylogenomic studies would therefore be promising to solve some of these questions.

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Research Article Mon, 2 Oct 2017 11:44:26 +0300
New findings of Flagellisargus J Zhang, 2012 (Diptera, Brachycera, Archisargidae), with discussion of the placements of some controversial taxa https://dez.pensoft.net/article/13550/ Deutsche Entomologische Zeitschrift 64(2): 111-122

DOI: 10.3897/dez.64.13550

Authors: Junfeng Zhang

Abstract: A new species of a new subgenus and a similar known species referred to the genus Flagellisargus J Zhang, 2012 are described and illustrated based on a male and a female impression fossils of these flies: Flagellisargus (Changbingisargus) parvussubgen. et sp. n. and Flagellisargus (Flagellisargus) cf. sinicus J Zhang, 2012. The latter taxon is the first record of a female Flagellisargus. Recently taken out of Archisargoidea, this study concludes that Flagellisargus should be an archisargid genus based on the known (male) and new (female) impression fossils. The placement of Daohugosargus J Zhang, 2012b is reassessed. It demonstrates close similarities in body structure and wing venation to archisargid flies, and can be retained as an archisargid genus. Archirhagio mostovskii J Zhang, 2015 is separated from Archirhagio zhangi K Zhang et al., 2009. Helempis Ren, 1998 could be, as a separate genus, placed in Archisarginae, Archisargidae.

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Research Article Thu, 3 Aug 2017 09:48:09 +0300
The Nevrorthidae, mistaken at all times: phylogeny and review of present knowledge (Holometabola, Neuropterida, Neuroptera) https://dez.pensoft.net/article/13028/ Deutsche Entomologische Zeitschrift 64(2): 77-110

DOI: 10.3897/dez.64.13028

Authors: Ulrike Aspöck, Horst Aspöck, Xingyue Liu

Abstract: This monographic review of the Nevrorthidae Nakahara, 1915, covers all 19 validly described, extant species worldwide that belong to one of the smallest families of the order Neuroptera. The family embraces four genera: Nevrorthus Costa, 1863 (with five species occurring in the Mediterranean region), Austroneurorthus Nakahara, 1958 (with two species restricted to eastern Australia), Nipponeurorthus Nakahara, 1958 (with 11 species from eastern Asia: Japanese islands, mainland China, Taiwan), and Sinoneurorthus Liu, H. Aspöck & U. Aspöck, 2012 (with one species recorded from mainland China). A comprehensive taxonomical treatment of all extant taxa is presented, including the scant available biological data. Distribution maps for all species are provided. A phylogenetic analysis based on morphological data from both extant and extinct taxa was performed. Austroneurorthus, together with Nevrorthus and some Eocene Baltic amber genera, form a monophylum. The disjunct distribution of modern nevrorthid genera demonstrates the relictual nature of the family and points to a historical biogeography that could have led to the formation of the present distribution pattern. Future discovery of fossil material might substantiate these claims. Sonnet for a Vulnerable Creature Is the Climate still fine? Still clean, the Riverine? Ruined rivulets run dry Fossils – tho’ living – may Die.

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Research Article Thu, 3 Aug 2017 09:09:46 +0300
Pseudochrysis Semenov, 1891 is the valid genus name for a group of cuckoo wasps frequently referred to as Pseudospinolia Linsenmaier, 1951 (Hymenoptera, Chrysididae) https://dez.pensoft.net/article/13005/ Deutsche Entomologische Zeitschrift 64(1): 69-75

DOI: 10.3897/dez.64.13005

Authors: Paolo Rosa, Maurizio Pavesi, Villu Soon, Oliver Niehuis

Abstract: The cuckoo wasp genus Pseudochrysis Semenov, 1891 is currently treated by several authors as a junior subjective synonym of Euchroeus Latreille, 1809, due to a type species designation by O. W. Richards in 1935. In the original description of the genus Pseudochrysis, Semenov (1891) distinguished two subordinated taxa within the genus Pseudochrysis: the subgenus Pseudochrysis and the subgenus Spintharis (sensu Dahlbom 1854). Semenov included three species in the subgenus Spintharis, but failed to mention any species included in the nominal subgenus. He was the first author, however, who listed in a subsequent publication (Semenov 1892) eleven species to be included in the nominal subgenus. According to the International Code of Zoological Nomenclature (ICZN 1999, Art. 67.2.2), these eleven species are deemed to have been listed in the original description. One of these, Chrysura humboldti Dahlbom, 1845, was explicitly designated by Semenov (1892) as type species of Pseudochrysis. We therefore consider the designation of Pseudochrysis (Spintharis) virgo Semenov, 1891 as type species of Pseudochrysis by Richards (1935) as invalid. The currently widely used genus name Pseudospinolia Linsenmaier, 1951 (type species Chrysis uniformis Dahlbom, 1854) is consequently to be regarded as a junior subjective synonym of Pseudochrysis, given the current circumscription of the genus Pseudospinolia (including both Pseudospinolia humboldti and Pseudospinolia uniformis).

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Short Communication Wed, 31 May 2017 10:07:27 +0300
On the enigmatic Sinonemestrius Hong & Wang, 1990, with description of a new species based on a complete fossil fly (Diptera, Brachycera, Tabanomorpha, Heterostomidae) https://dez.pensoft.net/article/11724/ Deutsche Entomologische Zeitschrift 64(1): 61-67

DOI: 10.3897/dez.64.11724

Authors: Junfeng Zhang

Abstract: A new species of Sinonemestrius Hong & Wang, 1990 is described and illustrated based on a complete compression fossil of the fly. Currently placed in Rhagionemestriidae or Xylophagidae, the present study concludes that the morphology of the new find indicates that Sinonemestrius is a heterostomid genus within Tabanomorpha. The placement of Ahirmoneura neimengguensis K-y Zhang et al., 2008 is reassessed. It demonstrates close similarities in body structure and wing venation to those of Sinonemestrius, and can be provisionally transferred from Nemestrinidae to Heterostomidae: Sinonemestriinae.

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Research Article Fri, 7 Apr 2017 11:31:15 +0300
Revisions of the Afrotropical genera of Argidae and species of Pampsilota Konow, 1899 (Hymenoptera, Tenthredinoidea) https://dez.pensoft.net/article/10800/ Deutsche Entomologische Zeitschrift 64(1): 1-25

DOI: 10.3897/dez.64.10800

Authors: Andrew D. Liston, Georg Goergen, Frank Koch

Abstract: The Afrotropical fauna contains five genera of Argidae. These are keyed. New subjective synonyms, followed by the valid name in brackets, are Calarge Enslin, 1911 [Arge Schrank, 1802], Calarge africana Enslin, 1911 [Arge congrua Konow, 1907], Clyparge Pasteels, 1963 [Scobina Lepeletier & Serville, 1828], Clyparge terminalis Pasteels, 1963 [Scobina poecila (Klug, 1834)], and Sterictophora [sic] afra Pasteels, 1963 [Sphacophilus afer comb. n., species inquirenda near S. monjarasi Smith & Morales-Reyes, 2015]. The type material of both C. terminalis and S. afra was probably collected in the New World, but labelled with the wrong locality “Kamerun”. An introduction of both species to Africa, not followed by long-term establishment, seems less likely. The removal of these taxa from the faunal list of the region is recommended. The nine known Afrotropical species of Pampsilota are revised, and an illustrated dichotomous identification key presented, with distribution maps for all species. Four species are here described as new to science: P. dahomeyanus Goergen, Koch & Liston, sp. n., P. nigeriae Liston & Koch, sp. n., P. tsavoensis Liston & Koch, sp. n., and P. zebra Liston & Koch, sp. n. Lectotypes are designated for Pampsilota afer Konow, 1899, and Cipdele africana Mocsáry, 1909. The immature stages and host plant of only one species are known: P. dahomeyanus on Lannea nigritana (Anacardiaceae). Its larval morphology strongly resembles that of European and North American species of Arge. We provisionally retain Pampsilota as a valid genus, although it could justifiably be treated as comprising merely a species group, or groups, within Arge.

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Research Article Thu, 19 Jan 2017 12:53:54 +0200
Hidden species within the genus Ocys Stephens: the widespread species O. harpaloides (Audinet-Serville) and O. tachysoides (Antoine) (Coleoptera, Carabidae, Bembidiini) https://dez.pensoft.net/article/10748/ Deutsche Entomologische Zeitschrift 63(2): 287-301

DOI: 10.3897/dez.63.10748

Authors: David R. Maddison, Roy Anderson

Abstract: Beetles previously considered to be Ocys harpaloides (Audinet-Serville) from northern Africa, Spain, France, the United Kingdom, France, and Belgium belong to two species. These species can be distinguished using DNA sequences of 28S rDNA, 18S rDNA, COI, CAD, and topoisomerase I. A key, diagnoses, and images are provided to enable identification of specimens based upon characteristics of male and female genitalia, as well as microsculpture and other external structures. Through examination of the holotype of Bembidium harpaloides v. tachysoides Antoine, as well as designation of lectotypes of Bembidion harpaloides Audinet-Serville and Ocys melanocephalus Stephens, and designation of a neotype for Tachis rufescens Guérin-Ménéville, the valid names of the two species were determined to be Ocys harpaloides and Ocys tachysoides (Antoine).

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Research Article Wed, 30 Nov 2016 10:17:09 +0200
Review of the world species of Exoryza (Hymenoptera, Braconidae, Microgastrinae), with description of five new species https://dez.pensoft.net/article/8977/ Deutsche Entomologische Zeitschrift 63(2): 195-210

DOI: 10.3897/dez.63.8977

Authors: Jose L. Fernandez-Triana, James B. Whitfield, M. Alex Smith, Tanya Dapkey, Winnie Hallwachs, Daniel H. Janzen

Abstract: The world species of the genus Exoryza (Hymenoptera, Braconidae, Microgastrinae) are revised. Ten species are recognized, including five new, authored by Fernandez-Triana: mariabustosae, richardashleyi, ritaashleyae, rosamatarritae and yeimycedenoae. The species Dolichogenidea oryzae Walker, 1994 is considered as related to Exoryza – although is not formally transferred to that genus. A dichotomous key to all species, descriptions and illustrations are provided. The only region where the genus is not yet recorded is Australasia, but this may be a collecting artifact. Host caterpillars (Lepidoptera) include species within Choreutidae, Depressariidae, Gelechiidae, and Pyraloidea – all but Pyraloidea are new host records. The status of Exoryza is questioned (it may only represent a species-group within the genus Dolichogenidea) but it is retained as a valid genus until a comprehensive phylogenetic study of Microgastrinae is available.

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Research Article Mon, 1 Aug 2016 11:41:01 +0300
Workers, soldiers, and gynes – morphometric characterization and description of the female castes of Camponotus singularis (Smith, 1858) (Hymenoptera, Formicidae) https://dez.pensoft.net/article/9435/ Deutsche Entomologische Zeitschrift 63(2): 183-193

DOI: 10.3897/dez.63.9435

Authors: Alice Laciny, Herbert Zettel, Irina Druzhinina

Abstract: Species of Camponotus Mayr, 1861 show a high variation of worker caste numbers, from a monomorphic worker caste to strong polymorphism along a continuous range of worker sizes or true worker dimorphism. Camponotus singularis (Smith, 1858) is used as a model for a Camponotus species with two very distinct worker castes (minors and majors) which are chiefly defined by morphometric data. We investigated shifts in proportions of C. singularis female castes in order to identify major allometric patters useful for characterizing caste differences in this species as well as in other Camponotini. We describe the main morphological traits which are characteristic for the respective morphs. The major worker, or “soldier”, shows many characteristics which deviate from the minor worker, but also from the alate gyne. Its traits are assumingly modified for its function in nest defence. Morphometric data clearly set Bornean specimens of Camponotus singularis (described as var. rufomaculatus Donisthorpe, 1941) apart from specimens with other proveniences, suggesting that this island population is a distinct species.

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Research Article Fri, 22 Jul 2016 15:02:20 +0300
Iuiuia caeca gen. n., sp. n., a new troglobitic planthopper in the family Kinnaridae (Hemiptera, Fulgoromorpha) from Brazil https://dez.pensoft.net/article/8432/ Deutsche Entomologische Zeitschrift 63(2): 171-181

DOI: 10.3897/dez.63.8432

Authors: Hannelore Hoch, Rodrigo Ferreira

Abstract: A new obligate cavernicolous (troglobitic) species in the planthopper family Kinnaridae is described from Brazil, and a new genus is established, as it could not be placed in any of the existing genera. Information on distribution and ecology is given. This is the second record of a troglobitic representative of this family from Brazil, and only the 6th cavernicolous kinnarid species worldwide.

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Research Article Mon, 6 Jun 2016 12:24:04 +0300
A review of the genus Xotidium Löbl, 1992 (Coleoptera, Staphylinidae, Scaphidiinae), with descriptions of five new species https://dez.pensoft.net/article/8386/ Deutsche Entomologische Zeitschrift 63(1): 155-169

DOI: 10.3897/dez.63.8386

Authors: Ryo Ogawa, Ivan Löbl

Abstract: In the present study, we have reviewed all species of the genus Xotidium Löbl, 1992, as well as five new species from Borneo, Malaysia, and Sulawesi, Indonesia, namely, Xotidium flagellum sp. n., X. heissi sp. n., X. meridionale sp. n., X. smetanai sp. n., X. tarantulatum sp. n. A key to all Xotidium species is provided. The female genitalic characters of Xotidium are illustrated and described for the first time.

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Research Article Tue, 31 May 2016 09:42:51 +0300
The Pselaphinae (Coleoptera, Staphylinidae) of New Caledonia and Loyalty Islands. III. Kieneriella, a new genus of the tribe Brachyglutini https://dez.pensoft.net/article/8344/ Deutsche Entomologische Zeitschrift 63(1): 149-154

DOI: 10.3897/dez.63.8344

Authors: Zi-Wei Yin, Peter Hlavac

Abstract: A new genus of the tribe Brachyglutini (Pselaphinae: Goniaceritae), Kieneriella gen. n., is described based on a new species K. novaecaledoniae sp. n. from New Caledonia. The strongly elongate maxillary palpi of Kieneriella is an unusual character state found in other genera of the tribe. The new taxon is compared with allied relatives, and its major diagnostic features are illustrated.

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Research Article Wed, 11 May 2016 10:00:11 +0300
Key to the species of Megarhyssa (Hymenoptera, Ichneumonidae, Rhyssinae) in America, north of Mexico https://dez.pensoft.net/article/7619/ Deutsche Entomologische Zeitschrift 63(1): 137-148

DOI: 10.3897/dez.63.7619

Authors: Victoria Pook, Michael Sharkey, David Wahl

Abstract: A dichotomous and an interactive key to the species of Megarhyssa (Hymenoptera: Ichneumonidae) in America, north of Mexico are presented. A diagnosis accompanied by images is provided for male and female wasps of each of the four species, Megarhyssa atrata, Megarhyssa greenei, Megarhyssa macrurus and Megarhyssa nortoni.

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Research Article Fri, 1 Apr 2016 09:27:29 +0300
Revision of the genus Trichrysis Lichtenstein, 1876 from China, with description of three new species (Hymenoptera, Chrysididae) https://dez.pensoft.net/article/7347/ Deutsche Entomologische Zeitschrift 63(1): 109-136

DOI: 10.3897/dez.63.7347

Authors: Paolo Rosa, Na-sen Wei, Jun Feng, Zai-fu Xu

Abstract: The Chinese species of the genus Trichrysis Lichtenstein, 1876 are revised for the first time. Thirteen species are recorded, of which three species are new for science, T. coeruleamaculata Rosa, Wei & Xu, sp. n., T. tridensnotata Rosa, Wei & Xu, sp. n., and T. yuani Rosa, Feng & Xu, sp. n.. Two species are revalidated: T. tonkinensis (Mocsáry, 1914) and T. formosana (Mocsáry, 1912). Two new synonymies are proposed, T. formosana (Mocsáry, 1912)=T. sauteri (Mocsáry, 1912), syn. n.=T. taial (Tsuneki, 1970), syn. n.. The lectotype of Chrysis pellucida du Buysson, 1887 is designated.

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Research Article Mon, 14 Mar 2016 09:05:50 +0200
Enigmatic distribution: first record of a hitherto New World planthopper taxon from Japan (Hemiptera, Fulgoroidea, Delphacidae, Plesiodelphacinae) https://dez.pensoft.net/article/7178/ Deutsche Entomologische Zeitschrift 63(1): 75-88

DOI: 10.3897/dez.63.7178

Authors: Manfred Asche, Masami Hayashi, Satoshi Fujinuma

Abstract: Burnilia japonica sp. n. of the delphacid subfamily Plesiodelphacinae from southern Japan (Kyushu, Yakushima, Okinawa) is described. The surprising discovery of a Burnilia-species in Japan is the first record of a member of this subfamily outside the New World. As the generic assignment is beyond any doubts, this finding reveals a puzzling geographic distribution of this group. A natural – indigenous – occurrence of B. japonica in Japan versus a recent introduction e.g., by human traffic, is discussed. A phylogenetic study of the whole Plesiodelphacinae including the Japanese species is desired.

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Research Article Thu, 10 Mar 2016 13:55:27 +0200
Species diversity, chorology, and biogeography of the Steninae MacLeay, 1825 of Iran, with comparative notes on Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae) https://dez.pensoft.net/article/5885/ Deutsche Entomologische Zeitschrift 63(1): 17-44

DOI: 10.3897/dez.63.5885

Authors: Sayeh Serri, Johannes Frisch

Abstract: The species diversity, chorology, and biogeography of the Steninae MacLeay, 1825 (Coleoptera: Staphylinidae) in Iran is described. A total of 68 species of Stenus Latreille, 1797 and one species of Dianous Leach, 1819 is recorded for this Middle Eastern country. Dianous coerulescens korgei Puthz, 2002, Stenus bicornis Puthz, 1972, S. butrintensis Smetana, 1959, S. cicindeloides Schaller, 1783, S. comma comma Le Conte, 1863, and S. hospes Erichson, 1840 are recorded for the Iranian fauna for the first time. Records of S. cordatoides Puthz, 1972, S. guttula P. Müller 1821, S. melanarius melanarius Stephens, 1833, S. planifrons planifrons Rey, 1884, S. pusillus Stephens, 1833, and S. umbricus Baudi di Selve, 1870 for Iran are, however, implausible or proved erroneous. Based on literature records and recent collecting data since 2004, the distribution of the stenine species in Iran is mapped, and their biogeographical relationships are discussed. As far as possible, the species are attributed to the commonly accepted distribution types in the Palaearctic Region after the theory of Pleistocene refuges of the arboreal biota. With the example of the Steninae and Scopaeus Erichson, 1839 (Staphylinidae: Paederinae), a cluster analysis is performed to examine the similarity of commonly accepted geographical units of Iran. The similar biogeographic pattern of the Steninae and Scopaeus reveal the high impact of Mediterranean elements on the fauna of Iran. The Hyrcanian subregion of the Caspian refuge is identified as the most significant center of origin of Stenus in Iran.

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Research Article Mon, 25 Jan 2016 11:06:19 +0200
Satrapister nitens Bickhardt, 1912: redescription and tentative phylogenetic placement of a mysterious taxon (Coleoptera, Histeridae, Saprininae) https://dez.pensoft.net/article/6363/ Deutsche Entomologische Zeitschrift 63(1): 1-8

DOI: 10.3897/dez.63.6363

Authors: Tomáš Lackner

Abstract: The monotypic genus Satrapister Bickhardt, 1912 is redescribed and figured. Its tentative position in the recently performed phylogeny of the subfamily, inferred from a new analysis based on the available morphological characters is discussed. Lectotype of Satrapister nitens Bickhardt, 1912 is designated.

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Research Article Fri, 8 Jan 2016 10:22:34 +0200
Descriptions of three new species and new records of Cerambycidae (Coleoptera) from America https://dez.pensoft.net/article/6615/ Deutsche Entomologische Zeitschrift 63(1): 9-16

DOI: 10.3897/dez.63.6615

Authors: Maria Helena M. Galileo, Larry G. Bezark, Antonio Santos-Silva

Abstract: Three new species are described and figured: Hemilissa birai sp. n. (Piezocerini), from Brazil; Estoloides (Estoloides) flavoscutellarissp. n. (Desmiphorini), from Ecuador; and Esthlogena (Esthlogena) setosasp. n. (Pteropliini), from Ecuador. Twenty-three new records in Lamiinae are provided.

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Review Article Fri, 8 Jan 2016 09:57:39 +0200
On foreign land: the conquest of Europe by Cinara curvipes (Patch, 1912) https://dez.pensoft.net/article/6457/ Deutsche Entomologische Zeitschrift 62(2): 261-265

DOI: 10.3897/dez.62.6457

Authors: Roman Hałaj, Barbara Osiadacz

Abstract: Cinara curvipes (Patch, 1912), a Nearctic aphid species first recorded in Europe at the turn of the millenium, has invaded almost one quarter of the continent during the past 15 years. This communication gives information about the first reports from Poland, summarises the species’ conquest of Europe and gives forecasts and possible explanations for such a rapid expansion. An identification key to aphid species on Abies is provided to help monitor its further expansion, as well as for the benefit of forestry and phytosanitary services.

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Short Communication Wed, 2 Dec 2015 08:49:15 +0200
‘The adikeshavus-group’: A new species group of Idris Förster (Hymenoptera, Platygastridae) from India, with descriptions of five new species https://dez.pensoft.net/article/6219/ Deutsche Entomologische Zeitschrift 62(2): 247-260

DOI: 10.3897/dez.62.6219

Authors: Veenakumari Kamalanathan, Prashanth Mohanraj, Farmanur Khan

Abstract: Idris Förster is a megagenus in the tribe Baeini comprising species that exclusively parasitize the eggs of spiders dwelling in vegetation and leaf litter. This is the only tribe in Platygastridae capable of using spider eggs for their development. Constructing species groups will facilitate studies of highly speciose genera like Idris. So far only one species group ‘the melleus-group’ has been proposed in this genus. A new species group ‘the adikeshavus-group’ from India is proposed. Five new species in this genus – I. adikeshavus, I. deergakombus, I. brevicornis, I. lopamudra and I. teestai – are described from India. All five species are imaged and a key to them is provided.

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Research Article Fri, 6 Nov 2015 15:21:38 +0200
Larval muscle attachment site (MAS) patterns are a conserved character among Piophilini flies (Diptera, Piophilidae) https://dez.pensoft.net/article/5685/ Deutsche Entomologische Zeitschrift 62(2): 239-245

DOI: 10.3897/dez.62.5685

Authors: Daniel Martín-Vega, Senta Niederegger

Abstract: The dorsoventral muscle attachment sites (MAS) patterns are described for six species of the tribe Piophilini (Diptera: Piophilidae): Centrophlebomyia furcata (Fabricius), Liopiophila varipes (Meigen), Piophila casei (Linnaeus), Piophila megastigmata McAlpine, Prochyliza nigrimana (Meigen) and Stearibia nigriceps (Meigen). Comparison between the MAS patterns of Piophilini and previous descriptions for Calliphoridae (Diptera) revealed differences in the muscle equipment between the larvae of both taxa. Among the Piophilini, the MAS patterns were highly conserved and only a genus-specific pattern for Piophila species and a species-specific pattern for C. furcata were found. Nevertheless, these differences in MAS patterns were subtle and some intraspecific variability was observed; hence, the MAS patterns do not appear to be suitable as diagnostic characters allowing for species determination of Piophilini larvae.

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Research Article Thu, 24 Sep 2015 18:34:04 +0300
A new genus and two new species of Pterostichini from China, with “sphodrine-like” parameres (Coleoptera, Carabidae) https://dez.pensoft.net/article/5493/ Deutsche Entomologische Zeitschrift 62(2): 225-237

DOI: 10.3897/dez.62.5493

Authors: Borislav Gueorguiev, Riccardo Sciaky

Abstract: A new genus of Pterostichini, Apophylon Guéorguiev & Sciaky, gen. n. (type species: Apophylon schillhammeri Guéorguiev & Sciaky, sp. n.) is proposed for Apophylon schillhammeri Guéorguiev & Sciaky, sp. n. (type locality: China, SE Guizhou Province, Leishan County, SE Kaili) and Apophylon pangu Guéorguiev & Sciaky, sp. n. (type locality: China, NW Hunan Province, Wulingyuan District, near the town of Wulingyuan). These two species share a unique combination of characters, not known in any other Pterostichini. The new genus can be distinguished by having (1) a left paramere without transverse apophysis; (2) a falcate right paramere, with styloid apex, broadened medial part and subbasal hasp; (3) a median lobe with dorsal ostium; (4) metatarsomeres 1-2 setose on the medioventral surface, in addition to the lateroventral setae; (5) meso- and metatarsomeres 1–4 with two dorsolateral grooves diverging distally; and (6) elytral striae with shining isodiametric microreticulation, which is in contrast to less shining transverse-mesh microreticulation on the elytral intervals.

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Research Article Thu, 3 Sep 2015 14:10:47 +0300
Contribution to the knowledge of the subgenus Scymnus (Parapullus) Yang, 1978 (Coleoptera, Coccinellidae), with description of eight new species https://dez.pensoft.net/article/5285/ Deutsche Entomologische Zeitschrift 62(2): 211-224

DOI: 10.3897/dez.62.5285

Authors: Xiaosheng Chen, Shunxiang Ren, Xingmin Wang

Abstract: Eight new species of the subgenus Scymnus (Parapullus) Yang, 1978 from China are described and illustrated: S. (P.) hastatussp. n., S. (P.) baxianshanensissp. n., S. (P.) laojielingensissp. n., S. (P.) annuliformissp. n., S. (P.) papillatussp. n., S. (P.) dichotomussp. n., S. (P.) shenmuensissp. n. and S. (P.) yanzigouensissp. n. Diagnoses and distributions are provided for each species. An updated key to the Chinese species of the subgenus Parapullus is given. A catalogue to all known species of this subgenus is also presented.

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Research Article Tue, 11 Aug 2015 12:07:54 +0300
New species, new synonymies and a new record of the genus Cryptogonus Mulsant, 1850 (Coleoptera, Coccinellidae) from China https://dez.pensoft.net/article/5242/ Deutsche Entomologische Zeitschrift 62(2): 203-210

DOI: 10.3897/dez.62.5242

Authors: Lizhi Huo, Wenjing Li, Xiaosheng Chen, Shunxiang Ren, Xingmin Wang

Abstract: Three new species of the genus Cryptogonus Mulsant, 1850 from China are described and illustrated: C. dulongjiangensis, C. fusiformis and C. reniformis Huo and Ren. Cryptogonus octoguttatus Mader, 1954 and C. kurosawai Sasaji, 1968 are recognized as synonymous with C. schraiki Mader, 1933. Cryptogonus hingstoni Kapur, 1948 is newly recorded from China. A species list of the genus Cryptogonus is presented.

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Research Article Tue, 11 Aug 2015 09:34:37 +0300
Revision of the genera Microplitis and Snellenius (Hymenoptera, Braconidae, Microgastrinae) from Area de Conservacion Guanacaste, Costa Rica, with a key to all species previously described from Mesoamerica https://dez.pensoft.net/article/5276/ Deutsche Entomologische Zeitschrift 62(2): 137-201

DOI: 10.3897/dez.62.5276

Authors: Jose L. Fernandez-Triana, James B. Whitfield, M. Alex Smith, Robert Kula, Winnie Hallwachs, Daniel Janzen

Abstract: The genera Microplitis and Snellenius (Hymenoptera: Braconidae, Microgastrinae) from Area de Conservacion Guanacaste (ACG), Costa Rica, are revised. A total of 28 new species are described: 23 of Snellenius (the first record for Mesoamerica) and five of Microplitis. A key is provided to all new species and five species of Microplitis previously described from Mesoamerica. In ACG, all Microplitis were reared exclusively from Sphingidae, while all Snellenius were reared from Noctuoidea (Noctuidae and Erebidae). All of the wasp species with known host records are unambiguously specialists, parasitizing one or a few related hosts. Biological information (wasp cocoon and caterpillar hosts) in the Neotropical region seems to differ from similar data reported in previous works for the Oriental region -but more studies on the world fauna are needed. Although the distinction between these two genera has been controversial, we consider that the available evidence, although not conclusive, suggests that these two genera are best kept as separate (based on the presence of at least a partial epicnemial carina in Snellenius, which is absent in Microplitis). The following 28 species, all authored by Fernández-Triana & Whitfield, are described as species nova: Microplitis adrianguadamuzi, M. alexanderrojasi, M. francopupulini, M. hebertbakeri, M. jorgehernandezi, Snellenius billburgeri, S. bobdressleri, S. donstonei, S. felipechavarriai, S. gerardoherrerai, S. irenebakerae, S. isidrochaconi, S. johnkressi, S. jorgecampabadali, S. jorgegomezlauritoi, S. josesarukhani, S. kerrydresslerae, S. lucindamcdadeae, S. luisdiegogomezi, S. mariakuzminae, S. mariamartachavarriae, S. phildevriesi, S. quiricojimenezi, S. robertoespinozai, S. sandyknappae, S. velvaruddae, S. vickifunkae, S. warrenwagneri.

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Research Article Mon, 20 Jul 2015 18:48:12 +0300
The Dilaridae of the Balkan Peninsula and of Anatolia (Insecta, Neuropterida, Neuroptera) https://dez.pensoft.net/article/5199/ Deutsche Entomologische Zeitschrift 62(2): 123-135

DOI: 10.3897/dez.62.5199

Authors: Ulrike Aspöck, Xingyue Liu, Horst Aspöck

Abstract: Basing upon all available information on type material of Dilar turcicus Hagen, 1858, Dilar syriacus Navás, 1909, and Dilar lineolatus Navás, 1909, together with a large number of dilarid specimens, the pleasing lacewings of Anatolia and Southeast Europe are revised. The current taxonomic concept of D. turcicus is confirmed, and a lectotype is designated. Dilar turcicus is widely distributed in the southeast of Europe (being the only representative of Dilaridae in this region), in Anatolia, and, most probably, in the Caucasus region. Dilar syriacus and D. lineolatus remain nomina dubia. Dilar syriacus might occur in Anatolia, while D. lineolatus is a species occurring in western Central Asia. Two new species, Dilar anatolicus sp. n. and Dilar fuscus sp. n. are described from Anatolia. Wings and genital segments of the three species occurring in Anatolia are illustrated, and a map documenting the known distribution of these species is provided.

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Research Article Fri, 3 Jul 2015 09:20:50 +0300
Contributions to the taxonomy of the ant genus Echinopla Smith, 1857 (Hymenoptera, Formicidae) https://dez.pensoft.net/article/5093/ Deutsche Entomologische Zeitschrift 62(1): 101-121

DOI: 10.3897/dez.62.5093

Authors: Herbert Zettel, Alice Laciny

Abstract: Sixteen species of Echinopla Smith, 1857 distributed west of Weber’s Line are treated, including eight species described as new: E. madli sp. n. from southern Thailand and Borneo, E. wardi sp. n. from West Malaysia, E. fisheri sp. n., E. circulus sp. n., E. mezgeri sp. n., and E. subtilis sp. n. from Borneo, E. brevisetosa sp. n. and E. angustata sp. n. from the Philippines. Lectotypes are designated for E. lineata Mayr, 1862 and E. senilis Mayr, 1862, stat. rev., and differences between the two species are pointed out. New records are presented for E. melanarctos Smith, 1857 (West Malaysia, Sumatra, Borneo), E. pallipes Smith, 1857 (Borneo), E. tritschleri Forel, 1901 (West Malaysia, Borneo), E. cherapunjiensis Bharti & Gul, 2012 (Laos, West Malaysia), E. lineata Mayr, 1862 (Singapore, Borneo), E. cf. pseudostriata Donisthorpe, 1943 (Philippines), and E. cf. vermiculata Emery, 1898 (Philippines).

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Research Article Thu, 4 Jun 2015 14:18:08 +0300
A revision of the genus Chlorocryptus Cameron (Hymenoptera, Ichneumonidae), with the first record of the genus from Japan https://dez.pensoft.net/article/5397/ Deutsche Entomologische Zeitschrift 62(1): 81-99

DOI: 10.3897/dez.62.8975

Authors: Takuma Yoshida, Rikio Matsumoto

Abstract: The genus Chlorocryptus is revised and two species are recognized: Chlorocryptus purpuratus (Smith, 1852) and C. coreanus (Szépligeti, 1916). Cryptaulax metallicus Szépligeti, 1916, which has been hitherto listed in catalogs as Chlorocryptus fuscipennis Townes et al., 1961, is shown as a new synonym of Chlorocryptus purpuratus. Chlorocryptus purpuratus and C. coreanus are redescribed and illustrated in detail and a key is provided. Chlorocryptus purpuratus is recorded from Japan and Vietnam for the first time and is considered to have recently been introduced into Japan.

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Research Article Tue, 2 Jun 2015 00:00:00 +0300
The Sawflies of Crete (Hymenoptera, Symphyta) https://dez.pensoft.net/article/4737/ Deutsche Entomologische Zeitschrift 62(1): 65-79

DOI: 10.3897/dez.62.4737

Authors: Andrew D. Liston, Hans-Joachim Jacobs, Marko Prous

Abstract: Forty-two sawfly species are now known from Crete, including twelve species here recorded for the first time, and excluding earlier published records of Allantus didymus (Klug, 1818) based on misidentifications. Allantus nigrolinearis (Zirngiebl, 1937) is treated as a distinct species within the laticinctus-didymus group. Aneugmenus oertzeni (Konow, 1887) is allied to the Sardo-Corsican A. bibolinii Zombori, 1979 and belongs to Aneugmenus Hartig, 1837 s. str., not Atoposelandria Enslin, 1913. Pristiphora nievesi Haris, 2002 is a new synonym of P. tetrica (Zaddach, 1883). Of the newly recorded species, Chevinia mediterranea Lacourt, 2003, Dolerus puncticollis Thomson, 1871 and Empria archangelskii Dovnar-Zapolskij, 1929 are simultaneously new for the country of Greece. Four species (Allantus nigrolinearis, Aneugmenus oertzeni, Periclista cretica (Schedl, 1981) and Pristiphora sp. [subbifida group]) have not been recorded outside Crete, and may be endemic. It is not clear whether the morphologically and genetically distinctive Cretan specimens of Strongylogaster multifasciata (Geoffroy, 1785) should be regarded as a Cretan endemic species, for which the name S. cretensis Konow, 1887 is available, or as an isolated population of S. multifasciata. This requires further study, as also the taxonomic status of Heterarthrus imbrosensis Schedl, 1981 (only known from Crete) and H. wuestneii (Konow, 1905) (widespread in the West Palaearctic). The sympatric occurrence of three related Pristiphora species on Acer sempervirens, two of which differ in their choice of host individuals that are at different stages of vegetative development, is remarkable. A checklist of the Symphyta of Crete is presented.

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Research Article Tue, 31 Mar 2015 15:49:48 +0300